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I'.ooK sK I.I.KK S THE ANIMAL KINGDOM, ARRANGED ACCORDING TO ITS ORGANIZATION, SERVING AS A FOUNDATION FOR THE NATURAL HISTORY OF ANIMALS, AND AN INTRODUCTION TO COMPARATIVE ANATOMY. BV BARON CUVIER, Great Officer of the Legion of Honour, Counsellor of State, and Member of the Royal Council of Public Instruction ; One of the Forty of the French Academy; Perpetual Secretary to the Academy of Sciences; Member of the Academies and Royal Societies of London, Berlin, Petersburgh, Stockholm, Turin, Edinburgh, Copenhagen, Gottingen, Bavaria, Modena, the Netherlands, and Calcutta ; and of the Linnsean Society of London, &c. &c. WITH FIGURES DESIGNED AFTER NATURE: THE BY M. LATREILLE, Chevalier of the Legion of Honour, Member of the Institute (Royal Academy of Sciences), and of the greater portion of other learned Societies in Europe, America, &c. ©ranslatrti from trje latest JFrcncI) 35t(ition. WITH ADDITIONAL NOTES, AND ILLUSTRATED BY NEARLY 500 ADDITIONAL PLATES. » IN FOUR VOLUMES. VOL. I. 230773 LONDON: G. HENDERSON, 2, OLD BAILEY, LUDGATE-HILL, AND SOLD BY ALL BOOKSELLERS. 1834. M'DOWALL. I-F.illBF.RTON ROW. COUGH SQUARK. i \& Lis- ADVERTISEMENT. IN presenting this version of the "Animal Kingdom" of the celebrated Cuvier to the British public, the Translator feels assured that he has only acted in compliance with the wishes of the most intelligent portion of the community, inasmuch as the great deficiency in our language of a complete work in this grand department of Natural History is thus supplied in a manner that it is impossible to excel. It is essential for the reader to understand that the attempts hitherto made by English authors to enrich British scientific literature with the labours of Cuvier, have been confined to the translation of the first edition of the " Regne Animale," which made its appearance so far back as the year 1816. With respect to that translation, it is not necessary that we should dwell upon it farther than to observe, that it is the version of a work which may now be deemed to be completely superseded. The great French author himself, indeed, has acknowledged the imperfections of his first edition, as compared with the last, which is now enriched with the results of labours, whereby, during the interval of twelve years, an immense pro- gress is declared by Cuvier to have been effected in this science. It is scarcely necessary to add, that no part of those labours, and no por- tion of that improvement, failed to be examined by this indefatigable naturalist. His connection with the government of France, his reputa- tion throughout Europe, and his consequent unbounded facilities of com- munication with fellow-labourers in all quarters of the globe, gave to Cuvier opportunities of procuring information of new facts, or corrections of former errors, such as could not be accessible to almost any other individual. From considering these facts, the reader will not fail to conclude that a difference, to no small amount, must necessarily exist between the former and the latter edition of the "Animal Kingdom;" nor will he, upon due examination, be prepared to deny that the latter is essentially a new and distinct work, from the number of alterations and improvements which have been incorporated with it. Cuvier records, with the most grateful expressions, his sense of the value of the information derived by him from the vast number of faithfully executed figures in Natural History which were supplied by recent travellers. The difficulties presented in the arrangement of the synonymes in the nomenclature of animals were also found by our great author very seriously diminished when he came to prepare the second edition of his " Regne Animale." Naturalists of [\ ADVERTISEMENT. all countries felt the necessity of more minute distinctions being esta- blished amongst those extensive groups which they had previously formed, and the result was a much nearer approximation than ever to an exact definition of each of the species. We refer to the Preface, at page xxx of the present volume, for a more copious account of the advantages which the last edition of the "Animal Kingdom" presents, as compared with the first. It remains for us, then, merely to state, that we felt the great importance of at once adding to our scientific literature a work of such permanent value as the " Animal Kingdom" of Cuvier. The character of the author for a profound knowledge of his subject — the conviction which we entertained of his exact accuracy in all that related to his labours, seemed to be sufficient to authorize us in trusting altogether to his authority ; and if we have added a few notes occasionally in the present volume, our object only was to enable our readers to make such an application of the text as our local advantages in this country enabled us to do. In the following work, therefore, the reader will not find himself di- verted from the regular current of the simple text by any protracted and tedious notes, which dispute, as it were, the right to the space of every page with the actual contents of the original. We have, in the front of our announcement to the public, pledged ourselves to place the British reader on a level with the French one, in comprehending the result of Cuvier's researches into the most interesting of the subjects that can engage the mind of man ; and to the fulfilment of that pledge we feel it to be our duty to adhere. Fidelity, then, in the translation, was the first grand object of our care. We have laid it down as a fixed rule, never to depart, even in a casual expression, from a most faithful representation of the thoughts and words of the original ; and we trust that we have not failed altogether in our attempt to transfer from his pages some portion at least of the energetic spirit, yet true simplicity, by which Cuvier's style is so happily distinguished. It has been, therefore, no object of our ambition, on this occasion, to attempt the improvement of the charming colours of the lily, or give fresh beauty to the glowing hues of the violet. Our task was plain ; and we felt that we performed enough, in devoting the whole of our exertions to effect the nearest possible approximation to the style and manner of the great Original — in other words, to secure to the English nation all the advantages of such an easy and instructive exposition of scientific knowledge as the French nation had already at their command. PREFACE TO THE FIRST EDITION. HAVING from ray earliest years devoted myself, from taste, to the study of Comparative Anatomy ; in other words, to the laws which preside over the organization of animals, and the modifications of that organization, as they are found throughout the diversified species — having, for nearly thirty years, consecrated to this science every moment which my duties left at my own disposal, I have ever kept in view, as the object of my labours, the resolution of the science into general laws, and into propositions of the simplest expression. My first essays soon made me perceive, that I could only attain this in proportion as the animals, whose structure I should have to elucidate, were arranged in conformity with that structure, so that in one single name of class, order, genus, &c. might be embraced all those species which, in their external as well as internal conformation, might have affinities either more general or particular. Now, this is what the greater number of naturalists of that epoch had never attempted, and what but few of them could have effected, had they even been willing to try, since a similar arrangement presupposes an extensive knowledge of the structures, of which it is partly the representation. It is true that Daubenton and Camper had supplied facts, — that Pallas had indicated views; but the ideas of these learned men had not yet exer- cised upon their contemporaries the influence which they merited. The only general catalogue of animals then in existence, and the only one we possess even now, the system of Linnaeus, had just been disfigured by an unfortunate editor, who did not even take the pains to examine the prin- ciples of that ingenious methodist, and who, wherever he found any dis- order, seems to have tried to render it more inextricable. It is no less true, that, upon particular classes, there existed some very extensive works, which described a considerable number of new species; but then the authors of these performances scarcely carried their attention beyond the external relations of these species, and no one was found to employ himself in arranging the classes and orders according to the nature of the structure of the animals. I was compelled then, — and the task occupied a considerable period of time, — to make anatomy and zoology, dissections and classification, the pioneers of my steps ; to search for better principles of distribution in my vot. r. c : viii PREFACE TO THE FIRST EDITION. first remarks on organization — to employ them in order to arrive at new ones, and to render the distribution perfect— in fine, from this mutual re- action of the two sciences, to elicit a system of zoology that might serve as an introduction and a guide in anatomical investigations, and as a body of anatomical doctrine fitted to develope and explain the zoological system. The first results of tins double labour appeared in 1795, in a special memoir upon a new division of the white-blooded animals. A sketch of their application to genera, and to their division in subgenera, was the object of my elementary " Tableau Elementaire des Animaux," printed in 1798, which, in conjunction with M. Dumeril, I improved, in the tables annexed to the first volumes of my " Lecons d'Anatomie Comparee" in 1800. I should, perhaps, have contented myself with perfecting these tables, and proceeded immediately to the publication of my great work on ana- tomy, if, in the course of my researches, I had not been frequently struck with another defect of the greater number of the general or partial sys- tems of zoology; I mean the confusion in which the want of critical acumen has left a great number of species, and even several genera. Not only were the classes and orders not in conformity with the in- timate nature of the animals, for the purpose of forming a foundation for a treatise on comparative anatomy; but the genera, though undoubtedly for the most part better composed, presented in their nomenclature very inadequate materials, inasmuch as the species were not arranged under each of them respectively according to its character. Thus, in placing the Sea- cow (Manatus, Cuv.) in the genus Morse (Trichechus, Lin.), the Siren in that of the Eels, Gmclin had rendered any general proposition relative to the organization of these two genera impossible, just as by approximating to the same class the same order, and placing side by side the Sepia and the fresh water Polypus, he had made it impossible to say any thing in general on the class and order which embraced such different beings. The examples above cited are selected from the most striking of these errors; but the number of them that existed was infinite, and, though not so easily to be perceived at the first glance, still they were not the less sources of real inconvenience. It was not enough, then, to have imagined a new arrangement of classes and orders, and to have properly placed the genera there; it was also ne- cessary to examine all the species, in order to ascertain if they really be- longed to the genera in which they had been placed. When I came to do this, I not only found that the species were either grouped or distributed in defiance of common sense ; but I saw that many of the species were by no means positively established by the characters attributed to them, or by the figures and descriptions given of them. PRKFACE TO TIIF, FfRST EDIT'ON. XiX 111 some parts, one of the species, by means of synonymes, is made to represent, under a single name, a great number, which are so different from each other as to be incapable of being placed in the same genus ; in others, a single species is doubled, and trebled, and appears again and again successively in divers subgenera, genera, and even sometimes in various orders. What shall we say, for instance, of the Trichechus Manatus of Gmelin, which in one single specific name comprises three species and two ge- nera; two genera, differing in almost every thing! By what name shall we speak of the Velella, which figures there twice among the Medusae, and once among the Holothurias? How are we to bring together the Biphorae; some of which are called there Dagysas, the greater number Salpae, and several placed among the Holothuriae? Thus, then, in order completely to attain the object, it was not sufficient to review the species — it was necessary to review even their synonymes, or, in other words, it was indispensable to reconstruct the system of animals. Such an enterprise, from the prodigious development of the science in late years, could not have been executed completely by any one individual, even supposing him to have no other employment, and to live the longest possible term of years. Had I been constrained to depend upon myself alone, I should not have been able to prepare even the simple sketch I now give ; but the resources of my position seemed to me to supply what I wanted both of time and talent. Living in the midst of so many able naturalists — drawing from their works as fast as they appeared — enjoying the use of their collections as freely as themselves — and having formed a very considerable one myself especially appropriated to my object, a great portion of my labour consisted merely in the employment of so many rich materials. It was not possible, for instance, that much remained for me to do on shells studied hy M. de Lamarck, or on quadrupeds described by M. Geoffroy. The numerous and new affinities observed by M. de Lacepede were so many traits for my system of fishes. Among so many beautiful birds, collected from all parts of the world, M. Le Vaillant per- ceived details of organization, which I immediately adapted to my plan. My own researches, employed and multiplied by other naturalists, yielded those fruits to me which, in my hands alone, they would not all have produced. Thus, by examining, in the cabinet I have formed, the anato- mical preparations on which I designed to found my division of reptiles. M. de Blainville and M. Oppel anticipated (and perhaps better than I could have done) results of which as yet I had but a glimpse, &c. &c. These reflections encouraged me; and I resolved on prefixing to my Treatise on Comparative Anatomy, a sort of abridged systematic table c 2 xx PREFACE TO THE FIRST EDITION. of animals, in which I should give their divisions and sub-divisions in the neatest detail, as established both in their internal and external structure ; in which I should indicate the best authenticated species be- longing to each of the subdivisions, and in which, to increase the interest, * should add some details regarding those species that are rendered re- markable by their being so common in this country, by their utility or mischievous practices, by the singularity of their habits and their economy, by their strange forms, their beauty, or their size. In so doing I hoped to prove useful to young naturalists, who, for the most part, have but little idea of the confusion and errors of criticism in which the most accredited works abound, and who, in foreign countries particularly, do not sufficiently attend to the study of the true relations of the conformation of beings; I considered myself as rendering a more direct service to those anatomists, who require to know beforehand to what orders they should direct their researches, when they wish to solve any problem of human anatomy or physiology by comparative anatomy, but whose ordinary occupa*ions do not sufficiently prepare them for ful- filling this condition, which is essential to their success. I had no intention, however, of extending this two-fold view to all the classes of the animal kingdom; and the Vertebrated animals, as in every sense the most interesting, naturally claimed a preference. Among the Invertebrata, I had to study more particularly the naked Mollusca and the great Zoophytes ; but the innumerable variations of the external forms of shells and corals, the microscopic animals, and the other families whose part, on the great theatre of nature, is not very apparent, or whose or- ganization affords but little room for the use of the scalpel, did not require a similar minuteness of detail. Independently of this, so far as the shells and corals were concerned, I could depend on the work of M. de Lamarck, in which will be found all that the most ardent thirst for knowledge can desire. As regards Insects, which, by their external form, organization, habits, and influence on all animated nature, are so highly interesting, I have been fortunate enough to find assistance, which, in rendering my work infinitely more perfect than it could have possibly been had it emanated from my pen alone, has, at the same time, considerably accelerated its publication. My friend and colleague, M. Latreille, who has studied these animals more profoundly than any other man in Europe, has kindly consented to give, in a single volume, and nearly in the order adopted for the other parts, a summary of his immense researches, and an abridged description of those innumerable genera entomologists are continually es- tablishing. With respect to the remaining portion, if I have given in some places PREFACE TO THE FIRST EDITION. XXI a less extended explanation of the subgenera and species, this imperfec- tion does not hold in the portion relating to the higher divisions and the relative characters, these being every where placed on foundations equally solid, the result of researches equally assiduous. I have examined, one by one, all the species of which I could procure specimens; I have approximated those which merely differed from each other in size, colour, or in the number of some parts of little importance, and have formed them into what I denominate subgenera. At every opportunity I dissected one species at least of each subgenus ; and if those be excepted to which the scalpel cannot be applied, there will then be but very few groups of this degree found in my work, of which I cannot produce some portion of the organs. Having determined the names of the species which I observed, and which had been previously either well described or well figured, I placed in the same subgenera those I had not seen, but whose exact figures, or descriptions, sufficiently precise to leave no doubt remaining as to their natural relations, I found in authors ; but I have passed over in silence that great number of vague indications, on which, in my opinion, natur- alists have been too eager to establish species, whose adoption is what has mainly contributed to introduce in the catalogue of animals that con- fusion which deprives it of so great a portion of its utility. 1 could, every where, have added great numbers of new species, but as I could not refer to figures it would in that case have been necessary to extend their descriptions beyond my limits; I have preferred, therefore, depriving my work of that ornament, and have indicated those only whose singular formation gives origin to new subgenera. My subgenera, once established on undoubted relations, and composed of well ascertained species, nothing remained but to construct this great scaffolding of genera, tribes, families, orders, classes and divisions which constitute the community of the animal kingdom. Here I have proceeded, partly by ascending from the inferior to the superior divisions, on the principles of affinity and comparison, and partly by descending from the superior to the inferior divisions, on the principle of the subordination of characters; carefully comparing the results of the two methods, verifying one by the other, and taking care to establish always the correspondence of forms, external and internal, both of which constitute integral parts of the essence of each animal. Such has been my mode of proceeding whenever it was necessary and possible to form new arrangements ; but I need not observe that, in many places, the results to which it would have conducted me had been already so satisfactorily obtained, that no other trouble was left to me than that of following the track of my predecessors. Even in these cases, how- ever, where I had nothing more to do than they had, by new observations XXII PREFACE TO THE FIRST EDITION. I have verified and confirmed what was previously acknowledged, and what I did not adopt until it was subjected to a rigorous scrutiny. An idea of this mode of examination may be obtained from the Memoirs on the anatomy of the Mollusca, which have appeared in the " Annales du Museum,'' and of which I am now preparing a separate and augmented collection. I venture to assure the reader, that the labour I have bestowed upon the Vertebrated animals, the Annulata, the Radiata, and many of the Insects and Crustacea, is equally extensive. I have not deemed it necessary to publish it with the same detail ; but all my preparations are exposed in the Cabinet of Comparative Anatomy in the Jardin du Roi, and will serve hereafter for my Treatise on Anatomy. Another work of considerable labour, but whose proofs cannot be made so authentic, is the critical examination of species. I have verified all the figures adduced by authors, and as often as possible referred each to its true species, before making a choice of those I have cited; it is from this verification alone, and never from the arrangement of preceding clas- sifiers, that I have referred to my subgenera the species that belong to them. Such is the reason why no astonishment should be experienced on finding that such or such a genus of Gmelin is now divided and dis- tributed even in different classes and divisions; that numerous nominal species are reduced to a single one, and that vulgar names are very dif ferently applied. There is not a single one of these changes that I an not prepared to justify, or of which the reader himself may not obtaii the proof by recurring to the sources I have indicated. In order to diminish this trouble, I have taken care to select for each class a principal author, generally the richest in good original figures, and I quote secondary works only in those cases in which the former are silent, or where it was useful to set up some comparison, for the sake ol better establishing synonymes. My subject could have been made to fill many volumes, but I consi- dered it my duty to condense it, by contriving abridged means of publi- cation. I have obtained these by graduated generalities; by never repeating for a species what could be said of a whole subgenus, nor for a genus what might be applied to an entire order, and thus is it that we ar- rive at the greatest possible economy of words. To this my endeavours have been, above all, particularly directed, inasmuch as this was the prin- cipal end of my work. It may be observed, however, that I have not employed many technical terms, and that 1 have endeavoured to commu- nicate my ideas without that barbarous apparatus of factitious words, which, in the works of so many modern naturalists, prove so very n pul- sive. I cannot perceive, however, that I have thereby lost any thing in precision or clearness. I have been compelled, unfortunately, to introduce many new names, PREFACE TO THE FIRST EDiTTON. XXtll although I endeavoured, as far as possible, to preserve those of my pre- decessors ; but the numerous subgenera I have established required these denominations, for in things so various the memory is not satisfied with numerical indications. I have selected them, either for an indication of some character, or from the common names which I have latinized, or finally, after the example of Linnaeus, from the mythological nomencla- ture, which are generally agreeable to the ear, and which we are far from having exhausted. In naming species, however, I would recommend the employment only of the substantive of the genus, and the trivial name. The names of the subgenera are designed as a mere relief to the memory, when we wish to indicate these subdivisions in particular. Otherwise, as the subgenera, already very numerous, will, in the end, become greatly multiplied, in consequence of having substantives continually to retain, we shall be in danger of losing the advantages of that binary nomenclature so happily imagined by Linnaeus. It is for the better preservation of it, that I have dismembered, as little as possible, the genera of that illustrious reformer of science. When- ever the subgenera in which I divide them were not to be translated to different families, I have left them together under their former generic appellation. This was not only due to the memory of Linneeus, but it was necessary in order to preserve the tradition and mutual understand- ing of the naturalists of different countries*. This habit, necessarily acquired in the study of natural history, of the mental classification of a great number of ideas, is one of the advantages of that science which is seldom observed, and which, when it shall have been generally introduced into the system of common education, will become, perhaps, the principal one. By it the student is exercised in that part of logic which is termed method, just as he is by geometry in that of syllo- gism, because natural history is the science which requires the most pre- cise methods, as geometry is that which demands the most rigorous rea- soning. Now this art of method, once well acquired, may be applied, with infinite advantage, to studies the most foreign to natural history. Every discussion implying a classification of facts, every inquiry which demands a distribution of materials, is performed according to the same laws; and the young man who had cultivated this science merely for * Here the author inserts a page of matter containing an explanation of the causes which induced him to direct the employment of several forms of type in the hody of the work, and also of the classes and other divisions which were to be indi- cated by the varieties of the letter. " Thus," he concludes the paragraph, " will the reader be able, at one glance, to distinguish the most important portions in every page, and the order of arrangement of every idea, and thus will the printer have se- conded the author in all those contrivances which his art is capable of supplying to the faculty of the memory." — End. Ed. XXIV PREFACE TO THE FIRST ED1I10N. amusement, is surprised, when lie makes the experiment, at the facilities it affords him in disentangling all kinds of affairs. It is not less useful in solitude. Sufficiently comprehensive to satisfy the most powerful mind, sufficiently various and interesting to calm the most agitated soul, it consoles the unhappy, and calms animosities. Once elevated to the contemplation of that harmony of nature irresistibly re- gulated by Providence, how weak and insignificant appear those causes which it has been pleased to leave dependent on the arbitrary will of man! How astonishing to behold so many examples of fine genius consuming themselves so vainly for their own happiness, or that of others, in the pur- suit of empty speculations, whose very traces a few years suffice to sweep away ! 1 boldly avow it — these ideas have always been present to my mind in my laborious hours; and if 1 have endeavoured by every means in my power to diffuse this peaceful study, it is because, in my opinion, it is more capable than any other of supplying that want of occupation which has so largely contributed to the disorders of our age — but I must return to my subject. There yet remains the task of accounting for the principal changes I have effected in the latest received methods, and to acknowledge the amount of my obligations to those naturalists who.se works have furnished or suggested a part of them. To anticipate a remark which will naturally present itself to many, I must observe that I have neither desired nor pretended to class animals so as to form one single line, or so as to mark their relative superiority. I even consider every attempt of this kind impracticable. Thus, I do not mean that such of the Mammalia or of the Birds as come last arc the most imperfect of their class ; still less do I believe that the last of the Mammalia are more perfect than the first of the Birds, the last of the Mollusca more so than the first of the Annulata or of the Radiata, even confining the meaning of this vague expression, must perfect, to that of most completely organized. I regard my divisions and subdivi- sions as merely the graduated expression of the resemblance of the beings which enter into each of them ; and although in some we observe a sort of degeneration or transition from one species to the other, which cannot be denied, this disposition is far from being general. The pre- tended scale of beings is but an erroneous application to the whole crea- tion of those partial observations, which are only true when confined to the limits within which they were made — and this application has, in my opinion, prejudiced the progress of natural history in modern times, to an extent which it is not easy to imagine. It is in conformity with these views that I have established my gi or Hemipodius, 319 Gen.- Subgen — Tinamus, — Turnix, 319 Syrrhaptes, 319 [Irypturus or Ynambus, 320 Subgen. — Pezus, 320 Tinamoos, 320 Gen.- — Columba, Rynchotus, 320 320 Subgen — Columbi-gallines, 321 Columba?, 321 Vinago, 323 V. GRALLATORI.E, 323 Fam. I. Brevipennes, 324 Gen. — Struthio, 324 Casuarius, 325 Fam. II. Pressirostres, 32G Gen.— Otis, 327 Charadrius, 327 Subgen. of Charadrius. — CEdicnemus, 327 Charadrius, 328 Gen. — Vanellus, 329 Subgen. — Vanellus, 329 INDEX. xIy V. GRALLATORIM— (continued). Gen. — Hematopus, 330 Cursorius or Tachydromus, 330 Cariama, Microdactyly, or Dicholophus, 331 Fam. III. Cultirostres, 331 Tribe 1.— Grus, 332 Gen.— Grus, 332 Subgen. — Psophia, 332 Grus, 333 Eurypyga, 333 Tribe 2 — Gen. — Cancroma, 334 Ardea, 334 Crab-eaters, 335 Onores, 335 Egrets, 335 Bitterns, 335 Night-herons, 336 Tribe 3.— Gen.— Ciconia, 336 Subgen. — Bare-necked Storks, 337 Pouched Storks, 337 Gen. — Mycteria, 337 Scopus, 338 Hians or Anastomus, 338 Subgen. — Droraas, 338 Gen. — Tantalus, 338 Platalea, 339 Fam. IV. Longirostres, 340 Gen. — Scolopax, 340 Subgen. — Ibis, 340 Numenius, 341 Scolopax, 342 Rhynchasa, 343 Limosa, 344 Calidris, 344 Arenaria, 345 Pelidna, 345 Cocorli, 345 Falcinellus, 346 Machetes, 346 Eurinorhynchus, 346 Phalaropus, 347 Strepsilas, 347 Totanus, 347 Lobipes, 349 Himantopus, 349 Gen. — Recurvirostra, 350 dvi INDEX. V. GRALLATORIJE— (continued). Fam. V. Macrodactyli: Gen. — Jacanas, 351 Palamedea, 352 Chauna, 352 Megapodius, 352 Rallus, 353 Fulica, 354 Subgen. of Fulica. — Gallinula, 354 Porphyrio, 354 Fulica, 355 Chionis or Vaginalis, 355 ~ t« . . oer Glareola, 355 Gen. — Phoenicopterus, 356 VI. PALMIPEDES, 357 Fam. J. Brachypter^:: Gen. — Colymbus : Subgen. — Podiceps, 358 Heliornis, 358 Mergus, 359 Uria, 359 Cephus, 360 Gen. — Alca, 360 Subgen. — Fractercula, 360 Alca, 361 Gen. — Aptenodytes, 361 Subgen. — Aptenodytes, 361 Catarrhactes, 362 Spheniscus, 362 Fam. II. Longipennes, 362 Gen. — Procellaria, 363 Subgen. — Puffinus, 364 Pelecanoides or Halodroma, 364 Pachyptila, 364 Gen. — Diomedea, 364 Larus, 365 Subgen. of Larus. — Goelands, 365 Mouettes, 366 Lestris, 366 Gen. — Sterna, 367 Subgen. — Noddies, 368 Gen. — Rhynchops, 368 Fam. III. Topipalmatje: Gen. — Pelecanus, 369 Subgen. — Pelecanus, 369 Phalacrocorax, 369 Tachypetes, 370 Sula, 370 Gen. — Plotus, 371 Phaeton, 371 index. xlvii VI. PALMIPEDES— (continued). Fam. IV. Lamellirostres. Gen. — Anas, 372 Subgen. — Cygnus, 372 Anser, 373 Barnacles, 374 Cereopsis, 374| Anas, 374 Oidemia, 375 Clangula, 375 Somateria, 376 Fuligula, 376 Rhynchapsis, 377 Tadorna, 378 Gen. — Mergus, 379 CORRIGENDA. In addition to the errors of haste of Cuvier, which we have noticed in the particular pages where they occur, there are others which we shall now point out. 234. By some unaccountable mistake, the Psaris Cuvieri of Swainson is confounded with the Pachy. semifasciatus of Spix. The first is as big as a sparrow, olive-green, with a yellow breast; the second is as big as a thrush, cream- coloured white, with black wings, tail, and crown. 235. for Sphecothere, read Sphecotherws. 238. for Sternura, read Stenura. — for T urdus volitans, read aurocapillus. 240. for M erremic, read Merrem., Ic. 249. for Cridotheres, read ^cridotheres. — note N.B. The genera Anthrochcera and Myzomcla, are not Swainson's, but those of Messrs. Horsfield and Vigors. 252. note N.B. The Oroolus regens is the Melliphaga chrysccephala (not regia) of Lewin, and the fyc. — The genus Tropedorhynchus is not Swainson's, but that of Horsfield and Vigors. 275. The Or. agripennis has been already noticed at p. 268 (under its original name of Emb. oryzevora), as the type of the genus Dolichonyx, Swainson. 284. note N.B. The genus Dasyornis is of Horsfield and Vigors; the rest following are Swainson's. The genera Peristera and Estopestes, among the Pigeons, are Swainson's. INTRODUCTION. j\S correct ideas respecting natural history are not very generally formed, it appears necessary to begin by defining its peculiar object, and establish- ing rigorous limits between it and neighbouring sciences. In our language, and in most others, the word nature is variously em- ployed. At one time it is used to express the qualities a being derives from birth, in opposition to those it may owe to art ; at another, the entire mass of beings which compose the universe; and at a third, the laws which govern those beings. It is in this latter sense particularly that we usually personify nature, and, through respect, use its name for that of its Creator. Physics, or Natural Philosophy, treats of the nature of these three re- lations, and is either general or particular. General physics examines abstractedly each of the properties of those moveable and extended beings we call bodies. That branch of them styled Dynamics, considers bodies in mass ; and, proceeding from a very small number of experiments, de- termines mathematically the laws of equilibrium, and those of motion and of its communication. Its different divisions are termed Statics, Hydro- statics, Hydrodynamics, Mechanics, &c. &c, according to the nature of the particular bodies whose motions it examines. Optics considers the particular motions of light, whose phenomena, which hitherto nothing but experiment has been able to determine, are becoming more numerous. Chemistry, another branch of general physics, exposes the laws by which the elementary molecules of bodies act on each other; the combinations or separations which result from the general tendency of these molecules to re-unite ; and the modifications which the various circumstances capable of separating or approximating them produce on that tendency. It is purely a science of experiment, and is irreducible to calculation. The theory of heat and that of electricity belong either to dynamics or chemistry, according to the point of view in which they are considered. VOL. I. A 2 INTRODUCTION. The ruling method in all the branches of general physics consists in isolating bodies, reducing them to their greatest simplicity, in bringing each of their properties separately into action, either by reflection or ex- periment, and by observing or calculating the results ; and finally, in gene- ralising and connecting the laws of these properties, so as to form codes, and, if it were possible, to refer them to one single principle into which they might all be resolved. The object of Particular Physics, or of Natural History — for the terms are synonymous — is, the special application of the laws recognised by the various branches of general physics to the numerous and varied beings which exist in nature, in order to explain the phenomena which each of them presents. Within this extensive range, Astronomy also would be included; but that science, sufficiently elucidated by mechanics, and completely sub- jected to its laws, employs methods, differing too widely from those re- quired by natural history, to permit it to be cultivated by the students of the latter. Natural history, then, is confined to objects which do not allow of exact calculation, nor of precise measurement in al! their parts. Meteorology also is subtracted from it and united to general physics ; so that, properly speaking, it considers only inanimate bodies called minerals, and the dif- ferent kinds of living beings, in all of which we may observe the effects, more or less various, of the laws of motion and chemical attraction, and of all the other causes analysed by general physics. Natural history, in strictness, should employ similar methods with the general sciences ; and it does so, in fact, whenever the objects it examines are sufficiently simple to allow it. This, however, is but very rarely the case. An essential difference between the general sciences and natural history is, that, in the former, phenomena are examined, whose conditions are all regulated by the examiner, in order, by their analysis, to arrive at general laws ; whereas, in the latter, they take place under circumstances beyond the control of him who studies them for the purpose of discovering, amid the complication, the effects of known general laws. He is not, like the experimenter, allowed to subtract them successively from each condition, and to reduce the problem to its elements — he is compelled to take it in its entireness, with all its conditions at once, and can perform the analysis only in thought. Suppose, for example, we attempt to insulate the nu- merous phenomena which compose the life of any of the higher orders of animals ; a single one being suppressed, every vestige of life is annihi- lated. Dynamics have thus nearly become a science of pure calculation ; che- Tti a INTRODUCTION. $ mistry is still a science of pure experiment ; and natural history, in a great number of its branches, will long remain one of pure observation. These three terms sufficiently designate the methods employed in the three branches of the natural sciences ; but in establishing between them very different degrees of certitude, they indicate, at the same time, the point to which they should incessantly tend, in order to attain nearer and nearer to perfection. Calculation, if we may so express it, thus commands nature, and deter- mines her phenomena more exactly than observation can make them known ; experiment compels her to unveil ; while observation pries into her secrets when refractory, and endeavours to surprise her. There is, however, a principle peculiar to natural history, which it uses with advantage on many occasions ; it is that of the conditions of existence, commonly styled final causes. As nothing can exist without the re-union of those conditions which render its existence possible, the component parts of each being must be so arranged as to render possible the whole being, not only with regard to itself but to its surrounding relations. The analysis of these conditions frequently conducts us to general laws, as cer- tain as those that are derived from calculation or experiment. It is only when all the laws of general physics and those which result from the conditions of existence are exhausted, that we are reduced to the simple laws of observation. The most effectual method of obtaining these, is that of comparison. This consists in successively observing the same bodies in the different positions in which nature places them, or in a mutual comparison of dif- ferent bodies ; until we have ascertained invariable relations between their structures and the phenomena they exhibit. These various bodies are kinds of experiments ready prepared by nature, who adds to or deducts from each of them different parts, just as we might wish to do in our labo- ratories ; shewing us, herself, at the same time their various results. In this way we finally succeed in establishing certain laws by which these relations are governed, and which are employed like those that are determined by the general sciences. The incorporation of these laws of observation with the general laws, either directly or by the principle of the conditions of existence, would complete the system of the natural sciences, in rendering sensible in all its parts the mutual influence of every being. To this end, should those who cultivate these sciences direct all their efforts. All researches of this nature, however, pre-suppose means of distin- guishing clearly, and causing others to distinguish, the bodies they are occupied with; otherwise we should be continually confounding them. Natural history then should be based on what is called a system of nature — • 4 INTRODUCTION. or, a great catalogue, in which all created beings have suitable names, may- be recognised by distinctive characters, and be arranged in divisions and subdivisions, themselves named and characterised, in which they may be found. In order that each being may be recognised in this catalogue, it must be accompanied by its character : habits or properties, which are but mo- mentary, cannot, then, furnish characters — they must be drawn from the conformation. There is scarcely a single being which has a simple character, or can be recognised by one single feature of its conformation ; a union of several of these traits are almost always required to distinguish one being from those that surround it, who also have some but not all of them, or who have them combined with others of which the first is destitute. The more numerous the beings to be distinguished, the greater should be the number of traits ; so that to distinguish an individual being from all others, a complete de- scription of it should enter into its character. It is to avoid this inconvenience, that divisions and subdivisions have been invented. A certain number only of neighbouring beings are com- pared with each other, and their characters need only to express their dif- ferences, which, by the supposition itself, are the least part of their con- formation. Such a re-union is termed a genus. The same inconvenience would be experienced in distinguishing genera from each other, were it not for the repetition of the operation in uniting the adjoining genera, so as to form an order, the orders to form a class, &c. Intermediate subdivisions may also be established. This scaffojding of divisions, the superior of which contain the inferior, is called a method. It is in some respects a sort of dictionary, in which we proceed from the properties of things to arrive at their names ; being the reverse of the common ones, in which we proceed from the name to arrive at the property. When the method is good, it does more than teach us names. If the subdivisions have not been established arbitrarily, but are based on the true fundamental relations, on the essential resemblances of beings, the method is the surest means of reducing the properties of beings to general rules, of expressing them in the fewest words, and of stamping them on the memory. To render it such, we employ an assiduous comparison of beings, di- rected by the principle of the subordination of characters, which is itself derived from that of the conditions of existence. The parts of a being pos- sessing a mutual adaptation, some traits of character exclude others, while, on the contrary, there are others that require them. When, therefore, we perceive such or such traits in a being, we can calculate before hand those that co-exist in it, or those that are incompatible with them. The parts, INTRODUCTION. 5 the properties, or the traits of conformation, which have the greatest num- ber of these relations of incompatibility or of co-existence with others, or, in other words, that exercise the most marked influence upon the whole of the being, are called the important characters, dominating characters ; the others are the subordinate characters, all varying in degree. This influence of characters is sometimes determined rationally, by the consideration of the nature of the organ. When this is impracticable, we have recourse to simple observation ; and a sure mark by which we may recognise the important characters, and one which is drawn from their own nature, is their superior constancy, and that in a long series of different beings, approximated according to their degrees of similitude, these cha- racters are the last to vary. That they should be preferred for distin- guishing the great divisions, and that, in proportion as we descend to the inferior subdivisions, we can also descend to subordinate and variable char- acters, is a rule resulting equally from their influence and constancy. There can be but one perfect method, which is the natural method. We thus name an arrangement in which beings of the same genus are placed nearer to each other than to those of the other genera; the genera of the same order nearer than those of the other orders, &c. &c. This method is the ideal to which natural history should tend ; for it is evident that if we can reach it, we shall have the exact and complete expression of all nature. In fact, each being is determined by its resemblance to others, and difference from them; and all these relations would be fully given by the arrangement in question. In a word, the natural method would be the whole science, and every step towards it tends to advance the science to perfection. Life being the most important of all the properties of beings, and the highest of all characters, it is not surprising that it has in all ages been made the most general principle of distinction; and that natural beings have always been separated into two immense divisions, the living and the inanimate. Of Living Beings, and Organization in general. If, in order to obtain a correct idea of the essence of life, we consider it in those beings in which its effects are the most simple, we quickly perceive that it consists in the faculty possessed by certain corporeal combinations, of continuing for a time and under a determinate form, by constantly at- tracting into their composition a part of surrounding substances, and ren- dering to the elements portions of their own. Life then is a vortex, more or less rapid, more or less complicated, the direction of which is invariable, and which always carries along molecules of similar kinds, but into which individual molecules are continually entering, 6 INTRODUCTION. and from which they are continually departing; so that the form of a living body is more essential to it than its matter. As long as this motion subsists, the body in which it takes place is living — it lives. When it finally ceases, it dies. After death, the ele- ments which compose it, abandoned to the ordinary chemical affinities, soon separate, from which, more or less quickly, results the dissolution of the once living body. It was then by the vital motion that its dissolution was arrested, and its elements were held in a temporary union. All living bodies die after a certain period, whose extreme limit is fixed for each species, and death appears to be a necessary consequence of life, which, by its own action, insensibly alters the structure of the body, so as to render its continuance impossible. In fact, the living body undergoes gradual, but continual changes, dur- ing the whole term of its existence. At first, it increases in dimensions, ac- cording to proportions, and within limits, fixed for each species and for each one of its parts; it then augments in density in the most of its parts — it is this second kind of change that appears to be the cause of natural death. If we examine the various living bodies more closely, we find they pos- sess a common structure, which a little reflection soon causes us to per- ceive is essential to a vortex, such as the vital motion. Solids, it is plain, are necessary to these bodies, for the maintenance of their forms ; and fluids for the conservation of motion in them. Their tissue, accordingly, is composed of network and plates, or of fibres and so- lid laminae, within whose interstices are contained the fluids ; it is in these fluids that the motion is most continued and extended. Foreign substan- ces penetrate the body and unite with them ; they nourish the solids by the interposition of their molecules, and also detach from them those that are superfluous. It is in a liquid or gaseous form that the matters to be exhaled traverse the pores of the living body ; but in return, it is the so- lids which contain the fluids, and by their contraction communicate to them part of their motion. This mutual action of the fluids and solids, this transition of molecules, required considerable affinity in their chemical composition ; and such is the fact — the solids of organized bodies being mostly composed of ele- ments easily convertible into fluids or gases. The motion of the fluids needing also a constantly repeated action on the parts of the solids, and communicating one to them, required in the latter both flexibility and dilatibility ; and accordingly we find this charac- ter nearly general in all organized solids. This structure, common to all living bodies ; this areolar tissue, whose more or less flexible fibres or laminae intercept fluids more or less abund- ant; constitutes what is called the organization. As a consequence of INTRODUCTION. 7 what we have said, it follows, that life can be enjoyed by organized bodies only. Organization, then, results from a great variety of arrangements, which are all conditions of life ; and it is easy to conceive, that if its effect be to alter either of these conditions, so as to arrest even one of the partial mo- tions of which it is composed, the general movement of life must cease. Every organized body, independently of the qualities common to its tis- sue, has a form peculiar to itself, not merely general and external, but ex- tending to the detail of the structure of each of its parts ; and it is upon this form, which determines the particular direction of each of the partial movements that take place in it, that depends the complication of the ge- neral movement of its life — it constitutes its species and renders it what it is. Each part co-operates in this general movement by a peculiar action, and experiences from it particular effects, so that in every being life is a whole, resulting from the mutual action and re-action of all its parts. Life, then, in general, pre-supposes organization in general, and the life proper to each individual being pre-supposes an organization peculiar to that being, just as the movement of a clock pre-supposes the clock; and accordingly we behold life only in beings that are organized and formed to enjoy it, and all the efforts of philosophy have never been able to discover matter in the act of organization, neither per se, nor by any external cause. In fact, life exercising upon the elements which at every moment form part of the living body, and upon those which it attracts to it, an ac- tion contrary to that which, without it, would be produced by the usual che- mical affinities, it seems impossible that it can be produced by these affini- ties, and yet we know of no other power in nature capable of re-uniting previously separated molecules. The birth of organized beings is, therefore, the greatest mystery of the organic economy and of all nature : we see them developed, but never be- ing formed; nay more, all those whose origin we can trace, have at first been attached to a body similar in form to their own, but which was deve- loped before them — in a word, to a parent. So long as the offspring has no independent existence, but participates in that of its parent, it is called a germ. The place to which the germ is attached, and the cause which detaches it, and gives it an independent life, vary; but this primitive adhesion to a similar being is a rule without exception. The separation of the germ is called generation. Every organized being reproduces others that are similar to itself, other- wise, death being a necessary consequence of life, the species would be- come extinct. Organized beings have even the faculty of reproducing, in degrees vary- 8 INTRODUCTION. ing with the species, particular parts of which they may have been de- prived — this is called the power of reproduction. The developement of organized beings is more or less rapid, and more or less extended, as circumstances are more or less favourable. Heat, the abundance and species of nutriment, with other causes, exercise great in- fluence, and this influence may extend to the whole body in general, or to certain organs in particular : thence arises the impossibility of a perfect similitude between the offspring and parent. Differences of this kind, between organized beings, form what are termed varieties. There is no proof, that all the differences, which now distinguish or- ganized beings, are such as may have been produced by circumstances. All that has been advanced upon this subject is hypothetical. Experi- ence, on the contrary, appears to prove, that, in the actual state of the globe, varieties are confined within rather narrow limits, and go back as far as we may, we still find those limits the same. We are thus compelled to admit of certain forms, which, from the origin of things, have perpetuated themselves without exceeding these limits, and every being appertaining to one or other of these forms constitutes what is termed a species. Varieties are accidental subdivisions of species. Generation being the only means of ascertaining the limits to which va- rieties may extend, species should be defined — the re-union of individuals descended one from the other, or from common parents, or from such as resemble them as strongly as they resemble each other. But although this definition is strict, it will be seen that its application to particular in- dividuals may be very difficult, where the necessary experiments have not been made. Thus then it stands — absorption, assimilation, exhalation, developement, and generation, are functions common to all living bodies ; birth and death the universal limits of their existence ; an areolar, contractile tissue, con- taining within its lamina fluids or gases in motion, the general essence of its structure ; substances, almost all susceptible of conversion into fluids or gases, and combinations capable of an easy and mutual transformation, the basis of their chemical composition. Fixed forms that are perpetuated by generation distinguish their species, determine the complication of the se- condary functions proper to each of them, and assign to them the parts they are to play on the great stage of the universe. These forms are nei- ther produced nor changed by their own agency — life supposes their exist- ence, its flame can only be kindled in an organization already prepared, and the most profound meditation and lynx-eyed and delicate observation can penetrate no farther than the mystery of the pre-existence of germs. INTRODUCTION. 9 Division of Organized Beings into Animals and Vegetables. Living or organized beings have always been subdivided into animate beings, that is, such as are possessed of sense and motion ; and into inani- mate beings, which are deprived of both these faculties, and are reduced to the simple faculty of vegetating. Although the leaves of several plants shrink from the touch, and the roots are steadily directed towards mois- ture, the leaves to light and air, and though parts of vegetables appear to oscillate without any apparent external cause, still these various motions have too little similarity to those of animals, to enable us to find in them any proofs of perception or will. The spontaneity in the motions of animals required essential modifications even in their purely vegetative organs. Their roots not penetrating the earth, it was necessary they should be able to place within themselves a supply of aliment, and to carry its reservoir along with them. Hence is derived the first character of animals, or their alimentary canal, from which their nutritive fluid penetrates all other parts through pores or vessels, which are a kind of internal roots. The organization of this cavity and its appurtenances required varying, according to the nature of the aliment, and the operation it had to under- go, before it could furnish juices fit for absorption; whilst the air and earth present to vegetables nought but elaborated juices ready for ab- sorption. The animal, whose functions are more numerous and varied than those of the plant, consequently necessitated an organization much more com- plete ; besides this, its parts not being capable of preserving one fixed re- lative position, there were no means by which external causes could pro- duce the motion of their fluids, which required an exemption from atmos- pheric influence ; from this originates the second character of animals — their circulating system, one less essential than that of digestion, since in the more simple animals it is unnecessary. The animal functions required organic systems not needed by vegetables — that of the muscles for volun- tary motion, and nerves for sensibility; and these two systems, like the rest, acting only through the motions and transformations of the fluids, it was necessary that these should be most numerous in animals, and that the chemical composition of the animal body be more complex than that of the plant ; and so it is, for one substance more (azote) enters into it as an es- sential element, whilst in plants it is a mere accidental junction with the three other general elements of organization — oxygen, hydrogen, and car- bon. This then is the third character of animals. From the sun and atmosphere, vegetables receive for their nutrition, water, which is composed of oxygen and hydrogen ; air, which contains 10 INTRODUCTION. oxygen and azote ; and carbonic acid, which is a combination of oxygen and carbon. To extract their own composition from these aliments, it was necessary they should retain the hydrogen and carbon, exhale the super- fluous oxygen, and absorb little or no azote. Such, in fact, is vegetable life, whose essential function is the exhalation of oxygen, which is effected through the agency of light. Animals also derive nourishment, directly or indirectly, from the vege- table itself, in which hydrogen and carbon form the principal parts. To assimilate them to their own composition, they must get rid of the su- perabundant hydrogen and carbon in particular, and accumulate more azote, which is performed through the medium of respiration, by which the oxygen of the atmosphere combines with the hydrogen and carbon ot their blood, and is exhaled with them in the form of water and carbonic acid. The azote, whatever part of the body it may penetrate, seems al- ways to remain there. The relations of vegetables and animals to the surrounding atmosphere are therefore in an inverse ratio — the former reject water and carbonic acid, while the latter produce them. The essential function of the animal body is respiration, it is that which in a manner annualizes it, and we shall see that the animal functions are the more completely exercised in proportion to the greatness of the powers of respiration possessed by the animal. This difference of relations constitutes the fourth character of animals. Of the Forms peculiar to the Organic Elements of the Animal Body, and of the principal Combinations of its Chemical Elements. An areolar tissue and three chemical elements are essential to every living body ; there is a fourth element peculiarly requisite to that of an animal ; but this tissue is composed of variously formed meshes, and these elements are variously combined. There are three kinds of organic materials or forms of texture — the cel- lular membrane, the muscular fibre, and the medullary matter; and to each form belongs a peculiar combination of chemical elements, as well as a particular function. The cellular substance is composed of an infinity of small fibres and lamina?, fortuitously disposed, so as to form little cells that communicate with each other. It is a kind of sponge, which has the same form as the body, all other parts of which traverse or fill it, and contracting indefi- nitely, on the removal of the causes of its tension. It is this power that retains the body in a given form and with certain limits. When condensed, this substance forms those lamina? called membranes ; the membranes rolled into cylinders, form those more or less ramified tubes INTRODUCTION. H named vessels; the filaments, called fibres, are resolved into it; and bones are nothing but the same thing indurated by the accumulation of earth- ly particles. The cellular substance consists of a combination well known as gelatine, characterised by its solubility in boiling water, and forming, when cold, a trembling jelly. We have not yet been able to reduce the medullary matter to its or- ganic molecules ; to the naked eye, it appears like a sort of soft bouillie, consisting of excessively small globules; it is not susceptible of any appa- rent motion, but in it resides the admirable power of transmitting to the me the impressions of the external senses, and conveying to the muscles the orders of the will. It constitutes the greater portion of the brain and the spinal marrow, and the nerves which are distributed to all the sen- tient organs are, essentially, mere fasciculi of its ramifications. The fleshy, or muscular fibre, is a peculiar sort of filament, whose distinc- tive property, during life, is that of contracting when touched or struck, or when it experiences the action of the will through the medium of the nerve. The muscles, direct organs of voluntary motion, are mere bundles of fleshy fibres. All vessels and membranes which have any kind of com- pression to execute are armed with these fibres. They are always inti- mately connected with nervous threads, but those which belong to the purely vegetative functions contract, without the knowledge of the me, so that, although the will is truly a means of causing the fibres to act, it is neither general nor unique. The fleshy fibre has for its base a particular substance called fibrine, which is insoluble in boiling water, and which seems naturally to assume this filamentous disposition. The nutritive fluid or the blood, such as we find it in the vessels of the circulation, is not only mostly resolvable into the general elements of the animal body, carbon, hydrogen, oxygen, and azote, but it also contains fi- brine and gelatine, almost prepared to contract and to assume the forms of membranes or filaments peculiar to them, all that is ever wanted for their manifestation being a little repose. The blood also contains another com- bination, which is found in many animal fluids and solids, called albumen, whose characteristic property is that of coagulating in boiling water. Be- sides these, the blood contains almost every element which may enter into the composition of the body of each animal, such as the lime and phospho- rus which harden the bones of vertebrated animals, the iron from which it and various other parts receive their colour, the fat or animal oil which is deposited in the cellular substance to supple it, &c. All the fluids and solids of the animal body are composed of chemical elements found in the 12 INTRODUCTION. blood, and it is only by possessing a few elements more or less, that each of them is distinguished ; whence it is plain, that their formation entirely depends on the subtraction of the whole or part of one or more elements of the blood, and in some few cases, on the addition of some element from elsewhere. These operations, by which the blood nourishes the fluid or solid matter of all parts of the body, may assume the general name of secretions. This name, however, is often appropriated exclusively to the production of li- quids ; while that of nutrition is more especially applied to the formation and deposition of the matter necessary to the growth and conservation of the solids. The composition of every solid organ, of every fluid, is precisely such as fits it for the part it is to play, and it preserves it as long as health re- mains, because the blood renews it as fast it becomes changed. The blood itself by this continued contribution is changed every moment, but is re- stored by digestion, which renews its matter by respiration, which delivers it from superfluous carbon and hydrogen, by perspiration and various other excretions, that relieve it from other superabundant principles. These perpetual changes of chemical composition form a part of the vi- tal vortex, not less essential than the visible movements and those of trans- lation. The object of the latter is, in fact, but to produce the former. Of the Forces which act in the Animal Body. The muscular fibre is not the only organ of voluntary motion, for we have just seen that it is also the most powerful of the agents employed by nature to produce those transmutations so necessary to vegetative life. Thus the fibres of the intestines produce the peristaltic motion, which causes the alimentary matter therein contained to pass through them ; the fibres of the heart and arteries are the agents of the circulation, and through it of all the secretions, &c. Volition contracts the fibre through the medium of the nerve ; and the involuntary fibres, such as those we have mentioned, being also animated by them, it is probable that these nerves are the cause of their con- traction. All contraction, and, generally speaking, every change of dimension in nature, is produced by a change of chemical composition, though it con- sist merely in the flowing or ebbing of an imponderable fluid, such as ca- loric; thus also are produced the most violent movements known upon earth, explosions, &c. There is, consequently, good reason to suppose that the nerve acts upon the fibre through the medium of an imponderable fluid, and the more so, as it is proved that this action is not mechanical. INTRODUCTION. 13 The medullary matter of the whole nervous system is homogeneous, and must be able to exercise its peculiar functions wherever it is found; all its ramifications are abundantly supplied with blood vessels. All the animal fluids being drawn from the blood by secretion, we can have no doubt that such is the case with the nervous fluid, and that the medullary matter secretes it. On the other hand, it is certain that the medullary matter is the sole conductor of the nervous fluid ; all the other organic elements restrain and arrest it, as glass arrests electricity. The external causes which are capable of producing sensations or caus ing contractions of the fibre are all chemical agents, capable of effecting decompositions, such as light, caloric, the salts, odorous vapours, percus- sion, compression, &rc. &c. It would appear then that these causes act on the nervous fluid chemi- cally, and by changing its composition ; this appears the more likely, as their action becomes weakened by continuance, as if the nervous fluid needed the resumption of its primitive composition to fit it for a fresh alteration. The external organs of the senses may be compared to sieves, which allow nothing to pass through to the nerve, except that species of agent which should affect it in that particular place, but which often accumulates it so as to increase its effect. The tongue has its spongy papillae which imbibe saline solutions ; the ear, a gelatinous pulp which is violently agi- tated by sonorous vibrations ; the eye, transparent lenses which concen- trate the rays of light, &c. &c. It is probable, that what are styled irritants, or the agents which occa- sion the contractions of the fibre, exert this action by producing on the fibre, by the nerve, a similar effect to that produced on it by the will; that is, by altering the nervous fluid, in the way that is requisite to change the dimensions of the fibre which it influences: but with this process the will has nothing to do, and very often the me is entirely ignorant of it. The muscles separated from the body preserve their susceptibility of irritation, as long as the portion of the nerve that remains with them preserves the power of acting on them — with this phenomenon the will has evidently no connexion. The nervous fluid is altered by muscular irritation, as well as by sen- sibility and voluntary motion, and the same necessity exists for the re- establishment of its primitive composition. The transmutations necessary to vegetable life are occasioned by irri- tants; the aliment irritates the intestine, the blood irritates the heart, &c. These movements are all independent of the will, and generally (while in health) take place without the knowledge of the me ; in several parts, the 14 INTRODUCTION. nerves that produce them are -even differently arranged from those that are appropriated to sensation, or dependent on the will, and the very object of this difference appears to be the securing of this independence. The nervous functions, that is, sensibility and muscular irritability, are so much the stronger at every point, in proportion as their exciting cause is abundant ; and as this cause, or the nervous fluid, is produced by secre- tion, its abundance must be in proportion to the quantity of medullary or secretory matter, and the amount of blood received by the latter. In animals that have a circulating system, the blood is propelled through the arteries which convey it to its destined parts, by means of their irrita- bility and that of the heart. If these arteries be irritated, they act more strongly, and propel a greater quantity of blood; the nervous fluid be- comes more abundant and augments the local sensibility ; this, in its turn, augments the irritability of the arteries, so that this mutual action may sometimes be carried to a great extent. It is called orgasm, and when it becomes painful and permanent, inflammation. The irritation may also originate in the nerve when exposed to the influence of acute sensations. This mutual influence of the nerves and fibres, either intestinal or ar- terial, is the real spring of vegetative life in animals. As each external sense is permeable only by such or such sensible sub- stances, so each internal organ may be accessible only to this or that agent of irritation. Thus, mercury irritates the salivary glands — cantharides irritate the bladder, &c. These agents are called specifics. The nervous system being homogeneous and continuous, local sensations and irritation debilitate the whole, and each function, by excessive action, may weaken the others. Excess of aliment weakens the power of thought, while long continued meditation impairs that of digestion, &c. Excessive local irritation will enfeeble the whole body, as if all the powers of life were concentrated in one single point. A second irritation produced at another part may diminish, or divert, as it is termed, the first : such is the effect of blisters, purgatives, &c. Brief as our sketch has been, it is sufficient to establish the possibility of accounting for all the phenomena of physical life, from the properties it presents, by the simple admission of a fluid such as we have defined. Summary Idea of the Functions and Organs of the Bodies of Animals, and of their various Degrees of Complication. After what we have stated respecting the organic elements of the body, its chemical principles and acting powers, nothing remains but to give a summary idea of the functions of which life is composed, and of their ap- propriate organs. INTRODUCTION. ] ') The functions of the animal body are divided into two classes : The animal functions, or those proper to animals, that is to say, sensi- bility and voluntary motion. The vital, vegetative functions, or those common to animals and vege- tables, i. e., nutrition and generation. Sensibility resides in the nervous system. The most general external sense is that of touch ; it is seated in the skin, a membrane that envelopes the whole body, which is traversed in every direction by nerves whose extreme filaments expand on the surface into papilla?, and are protected by the epidermis and other insensible tegu- ments, such as hairs, scales, &c. &c. Taste and smell are merely delicate states of the sense of touch, for which the skin of the mouth and nostrils is particularly organized: the first, by means of papillae more convex and spongy; the second, by its extreme delicacy and the multiplication of its ever humid surface. We have already spoken of the ear and the eye. The organ of generation is endowed with a sixth sense, seated in its in- ternal skin ; that of the stomach and intestines declares the state of those viscera by peculiar sensations. In fine, sensations more or less painful may originate in every part of the body through accident or disease. Many animals have neither ears nor nostrils, several are without eyes, and some are reduced to the single sense of touch, which is never absent. The action received by the external organs is continued by the nerves to the central masses of the nervous system, which, in the higher animals, consists of the brain and spinal marrow. The more elevated the nature of the animal, the more voluminous is the brain and the more is the sensitive power concentrated there ; the lower the animal, the more the medullary masses are dispersed, and in the most imperfect genera, the entire nervous substance seems to melt into the general matter of the body. That part of the body which contains the brain and principal organs of sense, is called the head. When the animal has received a sensation, and this has occasioned vo- lition, it is by the nerves, also, that this volition is transmitted to the muscles. The muscles are bundles of fleshy fibres whose contractions produce all the movements of the animal body. The extension of the limbs and every elongation, as well as every flexion and abbreviation of parts, are the ef- fects of muscular contraction. The muscles of every animal are arranged, both as respects number and direction, according to the movements it has to make ; and when these motions require force, the muscles are inserted into hard parts, articulated one over another, and may be considered as so many levers. These parts are called bones in the vertebrated animals, where they are internal, and are formed of a gelatinous mass, penetrated 16 INTRODUCTION. by particles of phosphate of lime. In the Mollusca, the Crustacea, and Insects, where they are external, and composed of a calcareous or horny substance that exudes between the skin and epidermis, they are called shells, crusts, and scales. The fleshy fibres are attached to the hard parts by means of other fibres of a gelatinous nature, which seem to be a continuation of the former, constituting what are called tendons. The configuration of the articulating surfaces of the hard parts limits their motion, which are also restrained by cords or envelopes, attached to the sides of the articulations, called ligaments. It is from the various arrangements of this bony and muscular appara- tus, and the form and proportion of the members therefrom resulting, that animals are capable of executing the innumerable movements that enter into walking and leaping, flight and natation. The muscular fibres, appropriated to digestion and the circulation, are independent of the will ; they receive nerves, however, but the chief of them are subdivided and arranged in a manner which seems to have for its object their independence of the me. It is only in paroxysms of the pas- sions and other powerful affections of the soul, which break down these barriers, that the empire of the me is perceptible, and even then it is al- most always to disorder these vegetative functions. It is, also, in a state of sickness only that these functions are accompanied with sensations : di- gestion is usually performed unconsciously. The aliment, divided by the jaws and teeth, or sucked up when liquids constitute the food, is swallowed by the muscular movements of the hinder parts of the mouth and throat, and deposited in the first portions of the ali- mentary canal that are usually expanded into one or more stomachs ; there it is penetrated with juices fitted to dissolve it. Passing thence through the rest of the canal, it receives other juices destined to complete its pre- paration. The parietes of the canal are pierced with pores which extract from this alimentary mass its nutritious portion; the useless residuum is rejected as excrement. The canal in which this first act of nutrition is performed, is a conti- nuation of the skin, and is composed of similar layers ; even the fibres that encircle it are analogous to those which adhere to the internal sur- face of the skin, called the fleshy pannicle. Throughout the whole inte- rior of this canal there is a transudation which has some connexion with the cutaneous perspiration, ard which becomes more abundant when the latter is suppressed ; the absorption of the skin is even very analogous to that of the intestines. It is only in the lowest order of animals that the ex- crements are rejected by the mouth, their intestines resembling a sac, having but one opening. INTRODUCTION. 17 Even among those where the intestinal canal has two orifices, there are many in which the nutritive juices, being absorbed by the parietes of the intestine, are immediately diffused throughout the whole spongy substance of the body : such, it would appear, is the case with all Insects. But from the Arachnides and Worms upwards, the nutritive fluid circulates in a system of closed vessels, whose ultimate ramifications alone dispense its molecules to the parts that are nourished by it; the vessels that convey it are called arteries, those that bring it back to the centre of the circulation, veins. The circulating vortex is here simple, and there double and even triple (including that of the vena-portce) ; the rapidity of its motion is often assisted by the contractions of a certain fleshy apparatus called a heart, which is placed at one or the other centres of circulation, and sometimes at both of them. In the red-blooded vertebrated animals, the nutritive fluid exudes from the intestines white or transparent, and is then termed chyle; it is poured into the veins, where it mingles with the blood, by a set of peculiar vessels called lacteals. Vessels similar to these lacteals, and forming with them an arrangement called the lymphatic system, also convey to the venous blood the residue of the nutrition of the parts and the products of cutaneous absorption. Before the blood is fit to nourish the parts, it must experience from the circumambient element the modification of which we have previously spoken. In animals possessing a circulating system, one portion of the vessels is destined to carry the blood into organs in which they spread it over a great surface to obtain an increase of this elemental influence. When that element is air, the surface is hollow, and is called lungs; when it is water, it is salient, and is termed branchice. There is always an ar- rangement of the organs cf motion for the purpose of propelling the ele- ment into, or upon, the organ of respiration. In animals destitute of a circulating system, air is diffused through every part of the body by elastic vessels called tracheae; or water acts upon them, either by penetrating through vessels, or by simply bathing the surface of the skin. The respired or purified blood is properly quali- fied for restoring the composition of all the parts, and to effect what is pro- perly called nutrition. This facility, which the blood possesses, of decom- posing itself at every point, so as to leave there the precise kind of mole- cule necessary, is indeed wonderful ; but it is this wonder which consti- tutes the whole vegetative life. For the nourishment of the solids we see no other arrangement than a great subdivision of the extreme arterial ra- mifications ; but for the production of fluids the apparatus is more complex and various. Sometimes the extremities of the vessels simply spread themselves over large surfaces, whence the produced fluid exhales; at vol. i. c 18 INTRODUCTION. others it oozes from the bottom of little cavities. Before these arterial extremities change into veins, they most commonly give rise to particular vessels that convey this fluid, which appears to proceed from the exact point of union between the two kinds of vessels ; in this case the blood vessels and these latter form, by interlacing, particular bodies called con- glomerate or secretory glands. In animals that have no circulation, in insects particularly, the parts are all bathed in the nutritive fluid: each of these parts draws from it what it requires, and if the production of a liquid be necessary, proper ves- sels floating in the fluid take up, by their pores, the constituent elements of that liquid. It is thus that the blood incessantly supports the composition of all the parts, and repairs the injuries arising from those changes which are the continual and necessary consequences of their functions. The general ideas we form with respect to this process are tolerably clear, although we have no distinct or detailed notion of what passes at each point ; and for want of knowing the chemical composition of each part with sufficient precision, we cannot render an exact account of the transmutations neces- sary to effect it. Besides the glands which separate from the blood those fluids that are destined for the internal economy, there are some which detach others from it that are to be totally ejected, either as superfluous — the urine, for instance, which is produced by the kidneys ; or for some use to the animal — as the ink of the cuttle-fish, and the purple matter of various mol- " lusca, &c. With respect to generation, there is a process or phenomenon, infinitely more difficult to comprehend than that of the secretions — the production of the germ. We have even seen that it is to be considered as almost in- comprehensible ; but the existence of the germ being admitted, generation presents no particular difficulties. As long as it adheres to the parent, it is nourished as if it were one of its organs ; and when it detaches itself, it possesses its own life, which is essentially similar to that of the adult. The germ, the embryo, the foetus, and the new-born animal have never, however, exactly the same form as the adult, and the difference is some- times so great, that their assimilation has been termed a metamorphosis. Thus, no one not previously aware of the fact, would suppose that the ca- terpillar is to become a butterfly. Every living being is more or less metamorphosed in the course of its growth; that is, it loses certain parts, and developes others. The anten- na?, wings, and all the parts of the butterfly were inclosed beneath the skin of the caterpillar ; this skin vanishes along with the jaws, feet, and other organs, that do not remain with the butterfly. The feet of the frog are INTRODUCTION. 19 inclosed by the skin of the tadpole; and the tadpole, to become a frog, parts with its tail, mouth, and branchiae. The child, at its birth, loses its placenta and membranes ; at a certain period its thymus gland nearly dis- appears, and it gradually acquires hair, teeth, and beard ; the relative size of its organs is altered, and its body augments in a greater ratio than its head, the head more than the internal ear, &c. The place where these germs are found, and the germs themselves are collectively styled the ovary; the canal through which, when detached, they are carried into the uterus, the oviduct; the cavity in which, in many species, they are compelled to remain for a longer or shorter period pre- vious to birth, the uterus; and the external orifice through which they pass into the world, the vulva. Where there are sexes, the male impreg- nates the germs appearing in the female. The fecundating liquor is called semen; the glands that separate it from the blood, testes; and when it is requisite it should be carried into the body of the female, the intro- ductory organ is named a penis. Of the Intellectual Functions of Animals. The impression of external objects upon the me, the production of a sensation or of an image, is a mystery into which the human understand- ing cannot penetrate ; and materialism an hypothesis, so much the more conjectural, as philosophy can furnish no direct proof of the actual exist- ence of matter. The naturalist, however, should examine what appear to be the material conditions of sensation, trace the ulterior operations of the mind, ascertain to w r hat point they reach in each being, and assure himself whether they are not subject to conditions of perfection, dependent on the organization of each species, or on the momentary state of each individual body. To enable the me to perceive, there must be an uninterrupted commu- nication between the external sense and the central masses of the medul- lary system. It is then the modification only experienced by these masses tbat the me perceives : there may also be real sensations, without the ex- ternal organ being affected, and which originate either in the nervous chain of communication, or in the central mass itself; such are dreams and vi- sions, or certain accidental sensations. By central masses, we mean a part of the nervous system, that is so much the more circumscribed, as the animal is more perfect. In man, it consists exclusively of a limited portion of the brain ; but in reptiles, it includes the brain and the whole of the medulla, and of each of their parts taken separately, so that the absence of the entire brain does not prevent sensation. In the inferior classes this extension is still greater. The perception acquired by the me, produces the image of the sensation c 2 20 INTRODUCTION. experienced. We trace to without the cause of that sensation, and thus acquire the idea of the object that has produced it. By a necessary law of our intelligence, all ideas of material objects are in time and space. The modifications experienced by the medullary masses leave impres- sions there which are reproduced, and thus recall to the mind images and ideas ; this is memory — a corporeal faculty that varies greatly, according to the age and health of the animal. Similar ideas, or such as have been acquired at the same time, recall each other ; this is the association of ideas. The order, extent, and quick- ness of this association constitute the perfection of memory. Every object presents itself to the memory with all its qualities or with all its accessary ideas. Intelligence has the power of separating these accessary ideas of objects, and of combining those that are alike in several different objects under a general idea; the object of which no where really exists, nor presents it- self per se — this is abstraction. Every sensation being more or less agreeable or disagreeable, experience and repeated essays soon shew what movements are required to procure the one and avoid the other ; and with respect to this, the intelligence abstracts itself from the general rules to direct the will. An agreeable sensation being liable to consequences that are not so, and vice versa, the subsequent sensations become associated with the idea of the primitive one, and modify the general rules framed by intelligence — this is prudence. From the application of these rules to general ideas, result certain for- mulae, which are afterwards easily adapted to particular cases — this is called reasoning. A lively remembrance of primitive and associated sensations, and of the impressions of pleasure or pain that belong to them, constitutes imagination. One privileged being, man, has the faculty of associating his general ideas with particular images more or less arbitrary, easily impressed upon the memory, and which serve to recall the general ideas they represent. These associated images are styled signs; their assemblage is a language. When the language is composed of images that relate to the sense of hear- ing, or of sounds, it is termed speech, and when relative to that of sight, hieroglyphics. Writing is a suite of images that relates to the sense of sight, by which we represent the elementary sounds, and by combining them, all the images relative to the sense of hearing of which speech is composed; it is therefore only a mediate representation of ideas. This faculty of representing general ideas by particular signs or images associated with them, enables us to retain distinctly, and to remember without embarrassment, an immense number; and furnishes to the rea- INTRODUCTION. 21 soiling faculty and the imagination innumerable materials, and to indivi- duals means of communication, which cause the whole species to participate in the experience of each individual, so that no bounds seem to be placed to the acquisition of knowledge — it is the distinguishing character of hu- man intelligence. Although, with respect to the intellectual faculties, the most perfect ani- mals are infinitely beneath man, it is certain that their intelligence per- forms operations of the same kind. They move in consequence of sensa- tions received, are susceptible of durable affections, and acquire by expe- rience a certain knowledge of things, by which they are governed inde- pendently of actual pain or pleasure, and by the simple foresight of con- sequences. When domesticated, they feel their subordination, know that the being who punishes them may refrain from so doing if he will, and, when sensible of having done wrong, or behold him angry, they assume a suppliant and deprecating air. In the society of man they become either corrupted or improved, and are susceptible of emulation and jealousy; they have among themselves a natural language, which, it is true, is merely the expression of their momentary sensations, but man teaches them to under- stand another, much more complicated, by which he makes known to them his will, and causes them to execute it. To sum up all, we perceive in the higher animals a certain degree of rea- son, with all its consequences, good and bad, and which appears to be about the same as that of children ere they have learned to speak. The lower we descend from man the weaker these faculties become, and at the bottom of the scale we find them reduced to signs (at times equivocal) of sensibi- lity, that is, to some few slight movements to escape from pain. Between these two extremes the degrees are infinite. In a great number of animals, however, there exists another kind of in- telligence, called instinct. This induces them to certain actions necessary to the preservation of the species, but very often altogether foreign to the apparent wants of the individual; often also very complicated, and which, if attributed to intelligence, would suppose a foresight and knowledge in the species that perform them, infinitely superior to what can possibly be granted. These actions, the result of instinct, are not the effect of imita- tion, for very frequently the individuals who execute them have never seen them performed by others : they are not proportioned to ordinary intelli- gence, but become more singular, more wise, more disinterested, in propor- tion as the annuals belong to less elevated classes, and in all the rest of their actions are more dull and stupid. They are so entirely the property of the species, that all its individuals perform them in the same way, with- out ever improving them a particle. The working bees, for instance, have always constructed very ingenious ?!? INTRODUCTION. edifices, agreeably to the rules of the highest geometry, and destined to lodge and nourish a posterity not even their own. The solitary bee, and the wasp also, form highly complicated nests, in which to deposit their eggs. From this egg comes a worm, which has never seen its parent, which is ig- norant of the structure of the prison in which it is confined, but which, once metamorphosed, constructs another precisely similar. The only method of obtaining a clear idea of instinct is by admitting the existence of innate and perpetual images or sensations in the sensorium, which cause the animal to act in the same way as ordinary or accidental sensations usually do. It is a kind of perpetual vision or dream that al- Avays pursues it, and it may be considered, in all that has relation to its instinct, as a kind of somnambulism. Instinct has been granted to animals as a supplement to intelligence, to concur with it, and with strength and fecundity, in the preservation, to a proper degree, of each species. There is no visible mark of instinct in the conformation of the animal, but, as well as it can be ascertained, the intelligence is always in propor- tion to the relative size of the brain, and particularly of its hemispheres. Of Method, as applied to the Animal Kingdom. From what has been stated with respect to methods in general, we have now to ascertain what are the essential characters in animals, on which their primary divisions are to be founded. It is evident that they should be those which are drawn from the animal functions, that is, from the sen- sations and motions ; for both these not only make the being an animal, but in a manner establish its degree of animality. Observation confirms this position by shewing that their degrees of de- velopement and complication accord with those of the organs of the vege- tative functions. The heart and the organs of the circulation form a kind of centre for the vegetative functions, as the brain and the trunk of the nervous system do for the animal ones. Now we see these two systems become imperfect and disappear together. In the lowest class of animals, where the nerves cease to be visible, the fibres are no longer distinct, and the organs of di- gestion are simple excavations in the homogeneous mass of the body. In insects the vascular system even disappears before the nervous one ; but, in general, the dispersion of the medullary masses accompanies that of the muscular agents : a spinal marrow, on which the knots or ganglions repre- sent so many brains, corresponds to a body divided into numerous rings, supported by pairs of limbs longitudinally distributed, &c. This correspondence of general forms, which results from the arrange- ment of the organs of motion, the distribution of the nervous masses, and INTRODUCTION. 23 the energy of the circulating system, should then be the basis of the pri- mary divisions of the animal kingdom. We will afterwards ascertain, in each of these divisions, what characters should succeed immediately to those, and form the basis of the primary subdivisions. General Distribution of the Animal Kingdom into four great Divisions. If, divesting ourselves of the prejudices founded on the divisions former- ly admitted, we consider only the organization and nature of animals, with- out regard to their size, utility, the greater or less knowledge we have of them, and other accessary circumstances, we shall find there are four prin- cipal forms — four general plans, if it may be so expressed, on which all ani- mals seem to have been modelled, and whose ulterior divisions, whatever be the titles with which naturalists have decorated them, are merely slight modifications, founded on the developement or addition of certain parts, which produce no essential change in the plan itself. In the first of these forms, which is that of man, and of the animals most nearly resembling him, the brain and principal trunk of the nervous system are inclosed in a bony envelope, formed by the cranium and vertebra? ; to the sides of this intermedial column are attached the ribs, and bones of the limbs, which form the frame work of the body ; the muscles generally cover the bones, whose motions they occasion, while the viscera are contained within the head and trunk. Animals of this form we shall denominate Animalia Vertebrata. They have all red blood, a muscular heart, a mouth furnished with two jaws situated either above or before each other, distinct organs of sight, hearing, smell, and taste placed in the cavities of the face, never more than four limbs, the sexes always separated, and a very similar distribution of the medullary masses and the principal branches of the nervous system. By a closer examination of each of the parts of this great series of ani- mals, we always discover some analogy, even in species the most remote from each other ; and may trace the gradations of one same plan from man to the last of the fishes. In the second form there is no skeleton ; the muscles are merely at- tached to the skin, which constitutes a soft contractile envelope, in which, in many species, are formed stony plates, called shells, whose position and production are analogous to those of the mucous body. The nervous sys- tem is contained within this general envelope along with the viscera, and is composed of several scattered masses connected by nervous filaments; the chief of these masses is placed on the oesophagus, and is called the brain. Of the four senses, the organs of two only are observable, those of taste and sight, the latter of which are even frequently wanting. One single 24 INTRODUCTION. family alone presents organs of hearing. There is always, however, a complete system of circulation, and particular organs for respiration. Those of digestion and secretion are nearly as complex as in the verte- brata. We will distinguish the animals of this second form by the ap- pellation of Animalia Mollusca. Although, as respects the external configuration of the parts, the gene- ral plan of their organization is not as uniform as that of the vertebrata ; there is always an equal degree of resemblance between them in the struc- ture and the functions. The third form is that remarked in worms, insects, &c. Their nervous system consists of two long cords, running longitudinally through the ab- domen, dilated at intervals into knots or ganglions. The first of these knots, placed over the oesophagus, and called brain, is scarcely any larger than those that are along the abdomen, with which they communicate by filaments that encircle the oesophagus like a necklace. The covering or envelope of the body is divided by transverse folds into a certain number of rings, whose teguments are sometimes soft, and sometimes hard ; the muscles, however, being always situated internally. Articulated limbs are frequently attached to the trunk; but very often there are none. We will call these animals Animalia Articulata, Or, articulated animals, in which is observed the transition from the cir- culation in closed vessels to nutrition by imbibition, and the corresponding one of respiration in circumscribed organs, to that effected by trachea? or air vessels distributed throughout the body. In them, the organs of taste and sight are the most distinct ; one single family alone presenting that of hearing. Their jaws, when they have any, are always lateral. The fourth form, which embraces all those animals known by the name of zoophytes, may also properly be denominated Animalia Radiata, Or, radiated animals. We have seen that the organs of sense and mo- tion in all the preceding ones are symmetrically arranged on the two sides of an axis. There is a posterior and anterior dissimilar face. In this last division, they are disposed like rays round a centre ; and this is the case even when they consist of but two series, for then the two faces are similar. They approximate to the homogeneity of plants, having no very distinct nervous system or particular organs of sense ; in some of them, it is even difficult to discover a vestige of circulation ; their respiratory or- gans are almost universally seated on the surface of the body, the intestine in the greater number is a mere sac without issue, and the lowest of the INTRODUCTION. 25 series are nothing but a sort of homogeneous pulp, endowed with motion and sensibility*. * Before my time, modern naturalists divided all invertebrated animals into two classes, Insects and Worms. I was the first who attacked this method; and in a memoir read before the Society of Natural History of Paris on the 10th of May, 1795, and printed in the Decade Philosophique, I presented a new division, in which I marked the characters and limits of the Mollusca, Crustacea, Insects and Worms, Echinodermata and Zoophytes. In a memoir read before the Institute on the 31st of December, 1801, I ascertained the red-blooded worms or Annelides. And finally, in a memoir read before the Institute in July, 1812, and printed in the Annales du Museum d'Histoire Naturelle, tome xix, I distributed these various classes in three divisions, each of which is analogous to a branch of the vertebrata. FIRST GREAT DIVISION THE ANIMAL KINGDOM. ANIMALIA VERTEBRATA. The bodies and limbs of vertebrated animals being supported by a frame- work or skeleton, composed of connected pieces that are moveable upon each other, their motions are certain and vigorous. The solidity of this support enables them to attain considerable size, and it is among them that the lar- gest animals are found. The great concentration of the nervous system, and the volume of its central portions, give energy and stability to their sentiments, whence re- sult superior intelligence and perfectibility. Their body always consists of a head, trunk, and members. The head is formed by the cranium which contains the brain, and by the face which is composed of two jaws, and of the receptacles of the senses. The trunk is supported by the spine and the ribs. The spine is formed of vertebra?, the first of which supports the head, that move upon each other, and are perforated by an annular opening, form- ing together a canal, in which is lodged that medullary production from which arise the nerves, called the spinal marrow. The spine, most commonly, is continued into a tail, extending beyond the posterior members. The ribs are a kind of semicircular hoops, which protect the sides of the cavity of the trunk, they are articulated at one extremity with the verte- bra?, and most generally at the other with the sternum ; sometimes, how- ever, they do not encircle the trunk, and there are genera in which they are hardly visible. There are never more than two pairs of members, but sometimes one or the other is wanting, or even both. Their forms vary according to the movements they have to execute. The superior members are converted into hands, feet, wings, or fins, and the inferior into feet or fins. The blood is always red, and appears to be so composed as to sustain a 2S ANIMALIA VERTEBRATA. peculiar energy of sentiment and muscular strength, but in various de- grees, corresponding to their quality of respiration : from which originates the subdivision of the Vertebrata into four classes. The external senses are always five in number, and reside in two eyes, two ears, two nostrds, the teguments of the tongue, and those of the body, generally. In some species, however, the eyes are obliterated. The nerves reach the medulla through the foramina of the vertebras or those of the cranium; they all seem to unite with this medulla, which, af- ter crossing its filaments, spreads out to form the various lobes of which the brain is composed, and terminates in the two medullary arches called hemispheres, whose volume is in proportion to the extent of the intelli- gence. There are always two jaws, the greatest motion is in the lower one, which rises and falls ; the upper jaw is sometimes immoveable. Both of these are almost always armed with teeth, excrescences of a peculiar na- ture, which in their chemical composition are very similar to that of bone, but which grow by layers and transudation ; one whole class, however, that of birds, has the jaws invested with horn, and the genus Testudo, in the class of reptiles, is in the same case. The intestinal canal traverses the body from the mouth to the anus, ex- periencing various enlargements and contractions, having appendages and receiving solvent fluids, one of which, the saliva, is discharged into the mouth. The others, which are poured into the intestine only, have vari- ous names : the two principal ones are — the juices of the gland called the pancreas, and bile, a product of another very large gland named the liver. While the digested aliment is traversing its canal, that portion of it which is fitted for nutrition, called the chyle, is absorbed by particular ves- sels styled lacteals, and carried into the veins ; the residue of the nourish- ment of the parts is also carried into the veins by vessels analogous to these lacteals, and forming with them one same system, called the lympha- tic system. The blood which has served to nourish the parts, and which has just been renewed by the chyle and lymph, is returned to the heart by the veins — but this blood is obliged, either wholly or in part, to pass into the organ of respiration, in order to regain its arterial nature, previous to being again sent through the system by the arteries. In the three first classes this respiratory organ consists of lungs, that is, a collection of cells into which air penetrates. In fish only, and in some reptiles, while young, it consists of branchiae or a series of lamina?, between which water passes. In all the Vertebrata, the blood which furnishes the liver with the mate- rials of the bile is venous blood, which has circulated partly in the parietes of the intestines, and partly in a peculiar body called the spleen, and ANIMALIA VERTEBRATA. 2f) wliicTi, after being united in a trunk called the vena-portae, is again sub- divided at the liver. All these animals have a particular secretion ; the urine, which is pro- duced hi two large glands, attached to the sides of the spine of the back, called kidneys — the liquid they secrete is most commonly poured into a reservoir, named bladder. The sexes are separate, and the female has always one or two ovaries, from which the eggs are detached at the instant of conception. The male fectKidifies them with the seminal fluid, but the mode varies greatly. In most of the genera of the three first classes, it requires an intromission of the fluid; in some reptiles, and in most of the fishes, it takes place after the exit of the egg. Subdivision of the Vertebrata into four Classes. We have just seen how far vertebrated animals resemble each other, they present, however, four great subdivisions or classes, characterised by the kind or power of their motions, which depend themselves on the quan- tity of their respiration, inasmuch as it is from this respiration that the muscular fibres derive the strength of their irritability. The quantity of respiration depends upon two agents : the first is the relative amount of blood which is poured into the respiratory organ in a given instant of time ; the second is the relative amount of oxygen which enters into the composition of the surrounding fluid. The quantity of the former depends upon the disposition of the organs of circulation and re- spiration. The organs of the circulation may be double, so that all the blood which is brought back from the various parts of the body by the veins, is forced to circulate through the respiratory organ, previous to resuming its former course through the arteries ; or they may be simple, so that a part only of the blood is obliged to pass through that organ, the remainder returning directly to the body. The latter is the case with reptiles. The quantity of their respiration, and all their qualities which depend on it, vary with the amount of blood thrown into the lungs at each pulsation. Fishes have a double circulation, but their organ of respiration is formed to execute its function through the medium of water; and their blood is only acted on by the portion of oxygen it contains, so that the quantity of their respiration is perhaps less than that of reptfles. In the mammalia the circulation is double, and the aerial respiration simple, that is, it is performed in the lungs only ; their quantity of respi- ration is, consequently, superior to that of reptiles, on account of the form ?,0 ANIMALIA VERTEBRATA. of their respiratory organ, and to that of fishes, from the nature of their surrounding element. The quantity of respiration in birds is even superior to that of quadru- peds, not only because they have a double circulation and an aerial respi- ration, but also because they respire by many other cavities besides the lungs, the air penetrating throughout their bodies, and bathing the branches of the aorta, as well as those of the pulmonary artery. Hence result the four different kinds of motion for which the four classes of vertebrated animals are more particularly designed: quadrupeds, in which the quantity of respiration is moderate, are generally formed to walk and run, both motions being characterized by precision and vigour; birds, which have more of it, possess the muscular strength and lightness requisite for flight ; reptiles, where it is diminished, are condemned to creep, and many of them pass a portion of their lives in a kind of torpor ; fishes, in fine, to execute their motions, require to be supported in a fluid whose specific gravity is nearly as great as their own. All the circumstances of organization peculiar to each of these four classes, and those especially which regard motion and the external sensa- tions, have a necessary relation with these essential characters. The mammalia, however, have particular characters in their viviparous mode of generation, in the manner by which the foetus is nourished in the uterus throiigh the medium of the placenta, and in the mammas by winch they suckle their young. The other classes, on the contrary, are oviparous, and if we compare them to the first, we shall find such numerous points of resemblance as announce a peculiar system of organization in the great general plan of the vertebrata. MAMMALIA. CLASS I. MAMMALIA. The mammalia are placed at the head of the animal kingdom, not only because it is the class to which man himself belongs, but also because it is that which enjoys the most numerous faculties, the most delicate sensa- tions, the most varied powers of motion, and in which all the different qualities seem combined in order to produce a more perfect degree of in- telligence, the one most fertile in resources, most susceptible of perfection, and least the slave of instinct. As their quantity of respiration is moderate, they are designed in gene- ral for walking on the earth, but with vigorous and continued steps. The forms of the articulations of their skeleton are, consequently, strictly de- fined, which determines all their motions with the most rigorous pre- cision. Some of them, however, by means of limbs considerably elongated, and extended membranes, raise themselves in the air; others have them so shortened, that they can move with facility in water only, though this does not deprive them of the general characters of the class. The upper jaw, in all of these animals, is fixed to the cranium ; the lower is formed of two pieces only, articulated by a projecting condyle to a fixed temporal bone : the neck consists of seven vertebrae, one single species excepted, which has nine ; the anterior ribs are attached before, by cartilage, to a sternum consisting of several vertical pieces ; their an- terior extremity commences in a shoulder-blade that is not articulated, but simply suspended in the flesh, often resting on the sternum by means of an intermediate bone, called a clavicle. This extremity is continued by an arm, a fore-arm, and a hand, the latter being composed of two ranges of small bones, called the carpus, of another range called the metacarpus, and of the fingers, each of which consists of two or three bones, termed phalanges. With the exception of the cetacea, the first part of the posterior extre- mity, in all animals of this class, is fixed to the spine, forming a girdle or pelvis, which, in youth, consists of three pairs of bones — the ilium which is attached to the spine, the pubis which forms the anterior part of the girdle, and the ischium, the posterior. At the point of union of these 32 MAMMALIA. three bones is situated the cavity with which the thigh is articulated, to which, in its turn, is attached the leg, formed of two bones, the tibia and fibula ; this extremity is terminated by parts similar to those of the hand, i. e. by a tarsus, metatarsus, and toes. The head of the mammalia is always articulated by two condyles, with the atlas, the first vertebra of the neck. The brain is always composed of two hemispheres, united by a medul- lary layer, called the corpus callosum, containing the ventricles, and en- veloping four pairs of tubercles, named the corpora striata, or striated bodies, the thalami nervorum opticorum, or beds of the optic nerves, and the nates and testes. Between the optic beds is a third ventricle, which communicates with a fourth under the cerebellum, the crura of which al- ways form a transverse prominence under the medulla oblongata, called the pons Varolii, or bridge of Varolius. The eye, invariably lodged in its orbit, is protected by two lids and a vestige of a third, and has its crystalline fixed by the ciliary processes — its sclerotic is simply cellular. The ear always contains a cavity called the tympanum, or drum, which communicates with the mouth by the Eustachian tube; the cavity itself is closed externally by a membrane called the membrana tympani, and con- tains a chain of four little bones, named the incus or anvil, malleus or hammer, the os orbiculare or circular bone, and the stapes or stirrup ; a vestibule, on the entrance of which rests the stapes, and which communi- cates with three semicircular canals ; and, finally, a cochlea, which ter- minates by one canal in the vestibule, and by the other in the tympanum. Their cranium is subdivided into three portions ; the anterior is formed by the two frontal and ethmoidal bones, the middle by the two ossa parie- talia and the os ethmoides, and the posterior by the os occipitis. Between the ossa parietalia, the sphenoidalis and the os occipitis, are interposed the two temporal bones, part of which belong properly to the face. In the foetus, the occipital bone is divided into four parts : the sphenoi- dal into two halves, which are again subdivided into three pairs of lateral wings; the temporal into three, one of which serves to complete the cra- nium, the second to close the labyrinth of the ear, the third to form the parietes of the tympanum, &c. These bony portions, still more numerous in the earliest period of the foetal existence, are united more or less promptly, according to the species, and the bones themselves finally be- come consolidated in the adult. Their face consists of the two maxillary bones, between which pass the nostrils; the two intermaxillaries are situated before, and the two ossa palati behind them ; between these descends the vomer, a bony process of the os ethmoides ; at the entrance of the nasal canal are placed the ossa MAMMALIA. o3 nasi; to its external parietes adhere the inferior turbinated bones, the superior ones which occupy its upper and posterior portion belonging to the os ethmoides. The jugal or cheek bone unites the maxillary to the temporal bone on each side, and frequently to the os frontis ; finally, the os unguis, and pars plana of the ethmoid bone occupy the internal angle of the orbit, and sometimes a part of the cheek. In the embryo state these bones also are much more subdivided. Their tongue is always fleshy, connected with a bone called the hyoides, which is composed of several pieces, and suspended from the cranium by ligaments. Their lungs, two in number, divided into lobes, and composed of an infinitude of cells, are always inclosed, without any adhesion, in a cavity formed by the ribs and diaphragm and lined by the pleura; the organ of voice is always at the upper extremity of the trachea; a fleshy curtain, called the velum palati, establishes a direct communication between their larynx and nasal canal. Their residence on the surface of the earth rendering them less exposed to the alternations of cold and heat, their tegument, the hair, is but mo- derately thick, and in such as inhabit warm climates even that is rare. The cetacea, which live exclusively in water, are the only ones that are altogether deprived of it. The abdominal cavity is lined with a membrane called the peritoneum, and the intestinal canal is suspended to a fold of it called the mesentery, which contains numerous conglobate glands in which the lacteals ramify : another production of the peritoneum, styled the epiploon, hangs in front of and under the intestines. The urine, which is retained for a time in the bladder, finds an exit in both sexes, with very few exceptions, by orifices in the organs of gene- ration. In all the mammalia, generation is essentially viviparous ; that is, the foetus, directly after conception, descends into the uterus enveloped in its membranes, the exterior of which is called chorion and the interior amnios; it fixes itself to the parietes of this cavity by one plexus or more of ves- sels called the placenta, which establishes a communication between it and the mother, by which it receives its nourishment, and most probably its oxygenation ; notwithstanding which, the foetus of the mammalia, at an early period, has a vesicle analogous to that which contains the yolk in the ovipara, receiving in like manner vessels from the mesentery. It has also another external bladder named the allantoid, which communicates with the urinary one by a canal called the urachus. Conception always requires an effectual coitus, in which the semen masculinum is thrown into the uterus of the female, VOL. I. D 34 MAMMALIA. The young are nourished for some time after birth by a fluid (milk) peculiar to animals of this class, which is produced by the mammae at the time .of parturition, and continues to be so as long as is necessary. It is from the mammce that this class derives its name ; and being a character peculiar to it, they distinguish it better than any other that is external*. Division of the Mammalia into Orders. The variable characters which form essential differences among the Mammalia are taken from the organs of touch, on which depends their degree of ability or address, and from the organs of manducation, which determine the nature of their aliment, and are all closely connected, not only with every thing relative to the function of digestion, but also with a multitude of other differences relating even to their intelligence. The degree of perfection of the organs of touch is estimated by the number and the pliability of the lingers, and from the greater or less ex- tent to which their extremities are enveloped by the nail or the hoof. A hoof, which completely envelopes the end of the toe, blunts its sensi- bility, and renders the foot incapable of seizing. The opposite extreme is when a nail, formed of one single lamina, covers oidy one of the faces of the extremity of the finger, leaving the other pos- sessed of all its delicacy. The nature of the food is known by the grinders, to the form of which the articulation of the jaws universally corresponds. To cut flesh, grinders are required as trenchant as a saw, and jaws fitted like scissars, having no other motion than a vertical one. For bruising roots or grains, flat-crowned grinders are necessary, and jaws that have a lateral motion; hi order that inequalities may always exist on the crown of these teeth, it is also requisite that their substance be composed of parts of unequal hardness, so that some may wear away faster than others. Hoofed animals are all necessarily herbivorous, and have flat-crowned grinders, inasmuch as their feet preclude the possibility of their seizing a living prey. Animals with unguiculated fingers were susceptible of more variety; their food is of all kinds ; and, independently of the form of their grinders, they differ greatly from each other in the pliability and delicacy of their fingers. There is one character with respect to this, which has immense influence on their dexterity, and greatly multiplies its powers ; it is the faculty of opposing the thumb to the finger for the purpose of seizing mi- * We shall find, however, in the sequel some doubts on this subject, arising from certain points in the family of the Monolremata. MAMMALIA. 3$ nute objects, constituting what is properly called a hand; a faculty which is carried to its highest perfection in man, in whom the whole anterior ex- tremity is free and capable of prehension. These various combinations, which strictly determine the nature of the different Mammalia, have given rise to the following orders : — Among the unguiculated animals, the first is Man, who, in addition to privileges of other descriptions, possesses hands at the anterior extremi- ties only, the posterior being designed to support him in an erect position. In the order ne^it to man, that of the quadrumana, we find hands at the four extremities. In another order, that of the carnaria, the thumb is not free, and cannot be opposed to the anterior extremities. Each of these orders has the three sorts of teeth, grinders, canini, and incisors or cutting teeth. In a fourth order, that of the rodentia, the toes differ but little from those of the Carnaria, but there are no canine teeth, and the incisors are placed in front of the mouth, and adapted to a very peculiar sort of man- ducation. Then come those animals whose toes are much cramped, and deeply sunk in large nails, which are generally curved; they have no incisors, and in some the canines disappear, while others have none of any descrip- tion. We comprise them all under the title of the Edentata. This distribution of the unguiculated animals would be perfect, and form a very regular series, were it not that New Holland has lately fur- nished us with a little collateral one, consisting of animals ivith pouches, the different genera of which are connected by a general similarity of or- ganization ; some of them, however, in the teeth and nature of their diet corresponding to the Carnaria, others to the Rodentia, and a third to the Edentata. The hoofed animals are less numerous, and have likewise fewer irregu- larities. The ruminantia, by their cloven foot, the absence of true incisors in their upper jaw, and their four stomachs, form an order that is very distinct. The remaining hoofed animals may all be united in a single order, which I shall call pachydermata or jumenta, the elephant excepted, which might constitute a separate one, and which is remotely connected with that of the Rodentia. In the last place, we find those of the Mammalia which have no poste- rior extremities, whose piscatory form and aquatic mode of life would in- duce us to form them into* a particular class, were it not that in every thing else their economy is similar to that in which we leave them. n 2 36 MAMMALIA. These are the hot-blooded fishes of the ancients, or the cetacea, which, uniting to the vigour of the other Mammalia the advantage of being sus- tained by the watery element, present to our wondering sight the most gigantic of animals. ORDER I. BIMANA. Man forms but one genus, and that genus the only one of its order. As his history is the more directly interesting to ourselves, and forms the point of comparison to which we refer that of other animals, we will speak of it more in detail. We will rapidly sketch every thing that is peculiar in each of his or- ganic systems, amidst all that he shares in common with other Mammalia; we will examine the advantages he derives from these peculiarities over other species ; we will describe the principal varieties of his race and their distinguishing characters, and finally point out the natural order in which his individual and social faculties are developed. Peculiar Conformation of Man. The foot of Man is very different from that of the Monkey ; it is large ; the leg bears vertically upon it ; the heel is expanded beneath ; the toes are short, and but slightly flexible ; the great toe, longer and larger than the rest, is placed on the same line with, and cannot be opposed to them. This foot, then, is peculiarly well adapted to support the body ; but cannot be used for seizing or climbing, and as the hands are not calculated for walking, Man is the only true bimanous and biped animal. The whole body of Man is arranged with a view to a vertical position. His feet, as just mentioned, furnish him with a base more extensive than that of any other of the Mammalia. The muscles which extend the foot and thigh are more vigorous, whence proceeds the projection of the calf and buttock ; the flexors of the leg are inserted higher up, which allows full extension of the knee, and renders the calf more apparent. The pelvis is wider, hence a greater separation of the thighs and feet, and that pyramidal form of the body so favourable to equilibrium. The necks of the thigh bones form an angle with the body of the bone, which increases still more the separation of the feet, and augments the basis of the body. Finally, the head in this vertical position is in equilibrium on the body, because its articulation is exactly under the middle of its mass. Where he to desire it, Man could not, with convenience, walk on all fours ; his short and nearly inflexible foot, and his long thigh, would bring the knee to the ground; his widely separated shoulders and his arms, too far extended from the median line, would ill support the upper portion of BIMANA. .>< his body. The great indented muscle, which, in quadrupeds, suspends, as in a girth, the body between the scapula?, is smaller in Man than in any one among them. The head is also heavier, both from the magnitude of the brain and the smallness of the sinuses or cavities of the bones; and yet the means of supporting it are weaker, for he has neither cervical ligament, nor are his vertebra? so arranged as to prevent their flexure forwards ; the result of this would be, that he could only keep his head in the same line with the spine, and then his eyes and mouth being directed towards the earth, he could not see before him ; — in the erect position, on the con- trary, the arrangement of these organs is every way perfect. The arteries which are sent to his brain, not being subdivided as in many quadrupeds, and the blood requisite for so voluminous an organ being carried into it with too much violence, frequent apoplexies would be the consequence of a horizontal position. Man, then, is formed for an erect position only. He thus preserves the entire use of his hands for the arts, while his organs of sense are most favourably situated for observation. These hands, which derive such advantages from their liberty, receive as many more from their structure. The thumb, longer in proportion than that of the Monkey, increases its facility of seizing small objects. All the fingers, the annularis excepted, have separate movements, a fa- culty possessed by no other animal, not even by the monkey. The nail, covering one side only of the extremity of the finger, acts as a support to the touch, without depriving it of an atom of its delicacy. The arms, to which these hands are attached, are strongly and firmly connected by the large scapula, the strong clavicle, &c. Man, so highly favoured as to dexterity, is not at all so with respect to force. His swiftness in running is greatly inferior to that of other animals of his size. Having neither projecting jaws, nor salient canine teeth, nor claws, he is destitute of offensive weapons ; and the sides and upper parts of his body being naked, unprovided even with hair, he is absolutely with- out defensive ones. Of all animals, he is also the longest in attaining the power necessary to provide for himself. This very weakness, however, is but one advantage more — it compels him to have recourse to that intelligence within, for which he is so emi- nently conspicuous. No quadruped approaches him in the magnitude and convolutions of the hemispheres of the brain, that is, in the part of this organ which is the principal instrument of the intellectual operations. The posterior portion of the same organ extends backwards, so as to form a second covering to the cerebellum : the very form of his cranium announces this magnitude of the brain, while the smallness of his face shews how slightly that por- tion of the nervous system which influences the external senses predomi- nates in him. These external sensations, moderate as they all are in Man, are never- theless extremely delicate and well balanced. His two eyes are directed forwards; he does not see on two sides at once, like many quadrupeds ; which produces more unity in the result of his sight, and concentrates his attention more closely on sensations of this kind. The ball and iris of his eye vary but little; this restrains the ac- tivity of his sight to a limited distance, and a determined degree of light. US MAMMALIA. His external ear, possessing but little mobility or extent, does not increase the intensity of sounds; and yet, of all animals, he best distinguishes the various degrees of intonation. His nostrils, more complicated than those of the monkey, are less so than those of all other genera; and yet he ap- pears to be the only animal whose sense of smell is sufficiently delicate to be affected by unpleasant odours. Delicacy of smell must have some in- fluence on that of taste ; and, independently of this, Man must have some advantage in this respect over other animals, those, at least, whose tongues are covered with scales. Lastly, the nicety of his tact results both from the delicacy of his teguments and the absence of all insensible parts, as well as from the form of his hand, which is better adapted than that of any other animal for suiting itself to every little superficial inequality. Man is pre-eminently distinguished in the organ of his voice ; of all the Mammalia, he alone possesses the faculty of articulating sounds, its pro- bable causes being the form of his mouth and the great mobility of his lips. From this results his most invaluable mode of communication ; for, of all the signs which can be conveniently employed for the transmission of ideas, variations of sound are those which can be perceived at the greatest dis- tance, and are the most extensive in their sphere of operation. The whole of his structure, even to the heart and great vessels, appears to have been framed with a view to a vertical position. The heart is placed obliquely on the diaphragm, and its point inclines to the left, thereby occasioning a distribution of the aorta, differing from that of most quad- rupeds. The natural food of man, judging from his structure, appears to consist of the fruits, roots, and other succulent parts of vegetables : his hands of- fer him every facility for gathering them ; his short, and but moderately strong jaws on the one hand, and his canini being equal in length to the remaining teeth, and his tubercular molares on the other, would allow him neither to feed on grass nor to devour flesh, were these aliments not pre- viously prepared by cooking. Once, however, possessed of fire, and those arts by which he is aided in seizing animals or killing them at a distance, every living being was rendered subservient to his nourishment, thereby giving him the means of an infinite multiplication of his species. His organs of digestion are in conformity with those of manducation; his stomach is simple, his intestinal canal of moderate length, the great in- testines well marked, his caecum short and thick and augmented by a small appendage, and his liver divided only into two large lobes and one small one ; his epiploon hangs in front of the intestines, and extends into the pelvis. To complete the hasty sketch of the anatomical structure of Man requisite for this introduction, we will add, that he has thirty-two vertebrae, of which seven belong to the neck, twelve to the back, five to the loins, five to the sacrum, and three to the coccyx. Seven pairs of his ribs are united with the sternum by elongated cartilages, and are called true ribs ; the five fol- lowing pairs are denominated false ones. His adult cranium is formed of eight bones; an occipitalis, two ossa temporis, two parietalia, and the frontal, ethmoidal and sphenoidal bones. The bones of his face are four- teen in number, two maxillaries, two ossa malae, each of which joins the temporal to the maxillary bone of its own side by a kind of handle called the zygomatic arch; two nasal bones, two ossa palati behind the palate, a vomer between the nostrils, two turbinated bones of the nose in the nos- B!MANA. 39 trils, two lachrymals (unguis) in the internal angles of the orbits and the single bone of the lower jaw. Each jaw has sixteen teeth; four cutting incisors in the middle, two pointed canines at the corners, and ten tuber - culated molares, five on each side. At the extremity of the spine of his scapula, is a tuberosity called the acromion, to which the clavicle is at- tached, and over its articulation is a point called the coracoid process, with which certain muscles are connected. The radius revolves upon the ulna, owing to the mode of its articulation with the humerus. The carpus has eight bones, four in each range ; the tarsus has seven ; those of the re- maining parts of the hand and foot may be easily counted by the number of fingers and toes. Enjoying uniform and regular supplies of nourishment, the fruit of his industry, Man is at all times inclined to the " plaisirs d'amour," without ever experiencing that irresistible and violent impetus which marks the passion in quadrupeds. His organ of generation is not upheld by a bony axis; the prepuce does not tie it down to the abdomen, and it hangs loosely in front of the pubis. Numerous and large veins which effect a rapid transfer of the blood of his testes to the general circulation, appear to contribute to the moderation of his desires. The uterus of woman is a simple oval cavity ; her mammas, only two in number, are placed upon her breast, and correspond with the facility she possesses of supporting her child upon her arm. Physical and Moral Decelopement of Man. The term of gestation in the human species is nine months; and but one child is usually produced at a birth, as in five hundred cases of partu- rition there is but one of twins ; more than the latter is extremely rare. The fcetus, a month old, is generally about one inch in height ; when two months, it is two inches and a half; when three, five inches ; in the fifth month, it is six or seven inches; in the seventh, it is eleven inches; in the eighth, fourteen, and in the ninth, eighteen inches. Those which are born prior to the seventh month usually die. The first or milk teeth begin to appear in a few months, commencing with the incisors. The number increases in two years to twenty, which, about the seventh year, are successively shed to make room for others. Of the twelve posterior molares which are permanent, there are four which make their appearance at four years and a half, and four at nine ; the last four are frequently not cut until the twentieth year. The growth of the fcetus is proportionably increased as it approaches the time of birth — that of the child, on the contrary, is always less and less. It has more than the fourth of its height when born ; it attains the half of it at two years and a half, and the three-fourths at nine or ten years ; its growth is completed about the eighteenth year. Man rarely exceeds the height of six feet, and as rarely remains under five. Woman is usually some inches shorter. Puberty is announced by external symptoms, from the tenth to the twelfth year in girls, and from the twelfth to the sixteenth in boys ; it ar- rives sooner in warm climates ; and neither sex (very rarely at least) is productive before or after that manifestation. Scarcely has the body gained the full period of its growth in height, before it begins to increase in bulk; fat accumulates in the cellular tissue, 10 MAMMALIA. the different vessels become gradually obstructed, the solids become rigid, and, after a life more or less long, more or less agitated, more or less painful, old age arrives with decrepitude, decay, and death. Man rarely lives beyond a hundred years, and most of the species, either from disease, accident, or old age, perish long before that term. The child needs the assistance of its mother much longer than her milk ; from this it obtains an education both moral and physical, and a mutual attachment is created that is fervent and durable. The nearly equal number of the two sexes, the difficulty of supporting more than one wife, when wealth does not supply the want of power, all go to prove that monogamy is the mode of union most natural to our species ; and as, wherever this kind of tie exists, the father participates in the education of his offspring, the length of time required for that education allows the birth of others — hence the natural permanence of the conjugal state. From the long period of infantile weakness springs domestic subordination, and the order of society in general, as the young people which compose the new families continue to preserve with their parents those tender re- lations to which they have so long been accustomed. This disposition to mutual assistance multiplies to an almost unlimited extent those advan- tages previously derived by insulated man from his intelligence; it has assisted him to tame or repulse other animals, to defend himself from the effects of climate, and thus enabled him to cover the earth with his species. In other respects, man appears to possess nothing resembling instinct, no regular habit of industry produced by innate ideas; his knowledge is the result of his sensations and of his observation, or of those of his prede- cessors. Transmitted by speech, increased by meditation, and applied to his necessities and his enjoyments, they have originated all the arts of life. Language and letters, by preserving acquired knowledge, are a source of indefinite perfection to his species. It is thus he has acquired ideas, and made all nature contribute to his wants. There are very different degrees of developement, however, in man. The first hordes, compelled to live by fishing and hunting, or on wild fruits, and being obliged to devote all their time .to search for the means of subsistence, and not being able to multiply greatly, because that would have destroyed the game, advanced but slowly. Their arts were limited to the construction of huts and canoes, to covering themselves with skins, and the fabrication of arrows and nets. They observed such stars only as directed them in their journeys, and some few natural objects whose pro- perties were of use to them. They domesticated the dog, simply because he had a natural inclination for their own kind of life. When they had succeeded in taming the herbivorous animals, they found in the possession of numerous flocks a never failing source of subsistence, and also some leisure, which they employed in extending the sphere of their acquire- ments. Some industry was then employed in the construction of dwell- ings and the making of clothes : the idea of property was admitted, and consequently that of barter, as well as wealth and difference of conditions, those fruitful sources of the noblest emulation and the vilest passions : but the necessity of searching for fresh pastures, and of obeying the changes of the seasons, still doomed them to a wandering life, and limited their improvement to a very narrow sphere, BIMANA. 41 The multiplication of the human species, and its improvement in the arts and sciences, have only been carried .to a high degree since the inven- tion of agriculture and the division of the soil into hereditary possessions. By means of agriculture, the manual labour of a portion of society is ade- quate to the maintenance of the whole, and allows the remainder time for less necessary occupations, at the same time that the hope of acquiring, by industry, a comfortable existence for self and posterity, has given a new spring to emulation. The discovery of a representative of property, or a circulating medium, by facilitating exchanges and rendering fortunes more independent and susceptible of being increased, has carried this emulation to its highest degree ; but, by a necessary consequence, it has also equally increased the vices of effeminacy and the furies of ambition. The natural propensity to reduce every thing to general principles, and to search for the causes of every phenomenon, has produced reflecting men, in every stage of society, who have added new ideas to those already obtained, nearly all of whom, while knowledge was confined to the few, endeavoured to convert their intellectual superiority into the means of do- mination, by exaggerating their own merit, and disguising the poverty of their knowledge by the propagation of superstitious ideas. An evil still more irremediable is the abuse of physical power: now that man only can injure man, he is continually seeking to do so, and is the only animal upon earth that is for ever at war with his own species. Savages fight for a forest, and herdsmen for a pasture, and, as often as they can, break in upon the cultivators of the earth to rob them of the fruits of their long and painful labours. Even civilized nations, far from being contented with their blessings, pour out each other's blood for the prero- gatives of pride, or the monopoly of trade. Hence, the necessity for go- vernments to direct the national wars, and to repress or reduce to regular forms the quarrels of individuals. The social condition of man has been restrained, or advanced by circum- stances more or less favourable. The glacial climates of the north of both continents, and the impenetra- ble forests of America, are still inhabited by the savage hunter or fisher- man. The immense sandy and salt plains of Central Asia and Africa are covered with a pastoral people and innumerable herds. These half civi- lized hordes assemble at the call of every enthusiastic chief, and rush like a torrent on the cultivated countries that surround them, in which they es- tablish themselves, but to be weakened by luxury, and in their turn to be- come the prey of others. This is the true cause of that despotism which has always crushed and destroyed the industry of Persia, India, and China. Mild climates, soils naturally irrigated and rich in vegetables, are the natural cradles of agriculture and civilization ; and when so situated as to be sheltered from the incursions of barbarians, every species of talent is excited; such were (the first in Europe) Greece and Italy, and such is, at present, nearly all that happy portion of the earth. There, are, however, certain intrinsic causes which seem to arrest the progress of particular races, although situated amidst the most favourable circumstances. 42 MAMMALIA. Varieties of the Human Species (a). Although the promiscuous intercourse of the human species, which pro- duces individuals capable of propagation, would seem to demonstrate its g^T (a) Notwithstanding the high character of Cuvier, as a founder of classes, yet the arrangement established by Blumenbach of the varieties of the human species has been universally adopted. In this classification the varieties are five, viz. — I. The Caucasian, which comprehends the ancient and modern inhabitants of Europe, the Western Asiatics, or those of this side of the Caspian Sea, and of the rivers Obi and the Ganges, together with the Northern Africans. The characters of this race are as follows: — The head is nearly the figure of a globe; the forehead is high and expanded; the cheek bones are without prominences; the nose is narrow and slightly aquiline; the face is oval and straight; the mouth small, with lips slightly everted; the skin is white, and the cheeks florid; the hair is long, soft, and shining, and varies in colour, from a nut-brown to the deepest black. — There are thirty-eight crania of this variety in the Hunterian Museum, London College of Surgeons. (See Plate I. Mammalia, Fig. 1. The portrait of Jusuf Aguiah Efendi, a Turk, and once Ambassador from the Sublime Porte at the Court of London). II. The Mongolian, commonly called the Tartarian, takes in the Finnish tribes in- habiting the colder parts of the north of Europe, such as the Laplanders and Esqui- maux, and also the Asiatics not included in the Caucasian variety, so that it com- prehends the Chinese, but not the Malays. The head approximates to a quadrilateral figure ; the face broad and flattened, so that the parts appear to run into each other ; the nose is small and flat, and the space between the eyes flat and broad; the cheek- bones are rounded and projecting; the aperture made by the eye-lids is narrow, and its line extends towards the temples, the internal angle of the eye being depressed towards the nose, and the upper eye-lid being at that angle continued into the lower one by a rounded sweep; the skin is pale olive, and the hair is thin, black, stiff", and straight. — There are nine crania of this variety in the Hunterian Museum. (See Plate I. Mammalia, Fig. 2. The portrait of Feodor Iwanowitsch, a Cal- muck, who was sent, when young, by the Empress of Russia to the Hereditary Princess of Baden; was educated at Carlsruhe, and became a famous engraver at Rome). III. The /Ethiopian, consists of all the Africans not included in the Caucasian divi- sion, and these partake more or less of the negro character. The front of the head is compressed laterally, and looks as if the forehead were removed, being, in this respect, a perfect contrast with the globular form of the head of the Caucasian va- riety. The entire cranium is contracted anteriorly, its cavity is considerably les- sened; the foramen magnum, and the condyles at its circumference, are placed farther back towards the occipital region; there is great developement of the face, and great prominence of the jaws, particularly of their alveolar margins and teeth, the upper incisors are oblique; the chin recedes, and the zygomatic arch projects to- wards the front; the skin is brown, black, and sometimes yellow, and the hair is deep black, crisp, and curly. — There are ten crania of this variety in the Hunterian Museum. (See Plate I. Mammalia, Fig, 3. The portrait of J. J. E. Capitein, a negro, who received holy orders in Holland). IV. The American, includes all the inhabitants of the vast continent of North and South America, excepting those of the northern part of the continent, and some of the islands, particularly the Caribbee. The cheeks are broad, but the malar bones are more rounded and arched than in the Mongolian race; the forehead is small and low; the orbits of the eye are unusually deep, and the nasal cavity is very large. The Caribs were in the habit of lowering the forehead by employing artificial pres- sure on the head in early infancy; hence, in this community, the characteristic feature of the American variety, the low forehead, is much more strikingly marked than in any other class of Americans. — There are five crania of this variety in the Hunterian Museum. (See Plate I. Mammalia, Fig. 4. The portrait of Thay Endaneega, a chief of the Mohawks, or Six Nations). V. The Malay, embraces the whole of the natives of the numerous Asiatic islands, and of those of the Pacific Ocean, New Zealand, New Holland, &c. Their head is BIMANA. 43 unity, certain hereditary peculiarities of conformation are observed, which constitute what are termed races. Three of them in particular appear very distinct — the Caucasian or white, the Mongolian or yellow, and the Ethiopian or negro. The Caucasian, to which we belong, is distinguished by the beauty of the oval formed by his head, varying in complexion and the colour of the hair. To this variety, the most highly civilized nations, and those which have generally held all others in subjection, are indebted for their origin. The Mongolian is known by his high cheek bones, flat visage, narrow and oblique eyes, straight black hair, scanty beard and olive complexion. Great empires have been established by this race in China and Japan, and their conquests been extended to this side of the Great Desert. In civi- lization, however, it has always remained stationary. The Negro race is confined to the south of mount Atlas; it is marked by a black complexion, crisped or woolly hair, compressed cranium, and a flat nose. The projection of the lower parts of the face, and the thick lips, evidently approximate it to the monkey tribe : the hordes of which it consists have always remained in the most complete state of utter bar- barism. The race from which we are descended has been called Caucasian, be- cause tradition and the filiation of nations seem to refer its origin to that group of mountains situated between the Caspian and Black seas, whence, as from a centre, it has been extended like the radii of a circle. Various nations in the vicinity of Caucasus, the Georgians and Circassians, are still considered the handsomest on earth. The principal ramifications of this race may be distinguished by the analogies of language. The Ar- menian or Syrian branch, stretching to the south, produced the Assyrians, the Chaldeans, the hitherto untameable Arabs, who, after Mahomet, were near becoming masters of the world; the Phoenicians, Jews, and Abyssini- ans, which were Arabian colonies ; and most probably the Egyptians. It is from this branch, always inclined to mysticism, that have sprung the most widely extended forms of religion — the arts and literature have some- times flourished among its nations, but always enveloped in a strange dis- guise and figurative style. The Indian, German, and Pelasgic branch is much more extended, and was much earlier divided: notwithstanding which, the most numerous affi- nities may be observed between its four principal languages — the Sanscrit, the present sacred language of the Hindoos, and the parent of the greater number of the dialects of Hindostan ; the ancient language of the Pelasgi, common mother of the Greek, Latin, many tongues that are extinct, and of all those of the south of Europe; the Gothic or Teutonic, from which are derived the languages of the north and north-west of Europe, such as the German, Dutch, English, Danish, Swedish, and other dialects; and moderately narrowed; the forehead is slightly arched; the face is large, and all its parts are fully developed; the jaws are more or less prominent; the skin is tawny, or clear mahogany or chesnut brown; the hair is black, soft, and curled. — There are thirty-four crania of this variety in the Hunterian Museum. (See Plate I. Mam- malia, Fig. 5. The portrait of Omai, a native of Ulietca, one of the Society Islands, brought to England in 1773, and carried back by Cook). — Eng. Ed. 44 MAMMALIA. finally, the Sclavonian, from which spring those of the north-east, the Rus- sian, Polish, Bohemian, &c. It is by this great and venerable branch of the Caucasian stock, that philosophy, the arts, and the sciences have been carried to the greatest perfection, and remained in the keeping of the nations which compose it for more than three thousand years. It was preceded in Europe by the Celts, who came from the north, whose tribes, once very numerous, are now confined to its most eastern extremity, and by the Cantabrians, who passed from Africa into Spain, now confounded with the many nations whose posterity have intermingled in that peninsula. The ancient Persians originate from the same source as the Indians, and their descendants to the present hour bear great marks of resemblance to the people of Europe. The predatory tribes of the Scythian and Tartar branch, extending at first to the north and north-east, always wandering over the immense plains of those countries, returned only to devastate the happier abodes of their more civilized brethren. The Scythians, who, at so remote a period, made irruptions into upper Asia; the Parthians, who there destroyed the Greek and Roman domination ; the Turks, who there subverted that of the Arabs, and subjugated in Europe the unfortunate remnant of the Grecian people, all swarmed from this prolific branch. The Finlanders and Hun- garians are tribes of the same division, which have strayed among the Sclavonic and Teutonic nations. Their original country, to the north and north-east of the Caspian sea still contains inhabitants who have the same origin, and speak similar languages, but mingled with other petty nations, variously descended, and of different languages. The Tartars remained unmixed longer than the others in the country included between the mouth of the Danube to beyond the Irtisch, from which they so long menaced Russia, and where they have finally been subjugated by her. The Mon- goles, however, have mingled their blood with that of those they con- quered, many traces of which may still be found among the inhabitants of lesser Tartary. It is to the east of this Tartar branch of the Caucasian race that the Mongolian race begins, whence it extends to the eastern ocean. Its branches, the Calmucs, &c, still wandering shepherds, are constantly tra- versing the desert. Thrice did their ancestors, under Attila, Genghis, and Tamerlane, spread far the terror of their name. The Chinese are the earliest and most civilized branch, not only of this race, to which they belong, but of all the nations upon earth. A third branch, the Mant- chures, recently conquered and still govern China. The Japanese, Co- reans, and nearly all the hordes which extend to the north-east of Siberia, subject to Russia, are also to be considered, in a great measure, as ori- ginating from this race ; and such also is esteemed the fact, with regard to the original inhabitants of various islands of that Archipelago. With the exception of a few Chinese literati, the different nations of the Mongoles are universally addicted to Buddism, or the religion of Fo. The origin of this great race appears to have been in the mountains of Atlai, but it is impossible to trace the filiation of its different branches with the same certainty as we have done those of the Caucasian. The history of these wandering nations is as fugitive as their establishments; BlMANA. 45 and that of the Chinese, confined exclusively to their own empire, gives us nothing satisfactory with respect to their neighbours. The affinities of their languages are also too little known to direct us in this labyrinth. The languages of the north of the Peninsula beyond the Ganges, as well as that of Thibet, are somewhat allied to the Chinese, at least in their monosyllabic structure, and the people who speak them have features somewhat resembling other Mongoles. The south of this Peninsula, how- ever, is inhabited by Malays, whose forms approximate them much nearer to the Indians, whose race and language are extended over all the coasts of the islands of the Indian Archipelago. The innumerable little islands of the southern ocean are also peopled by a handsome race, nearly allied to the Indians, whose language is very similar to the Malay; in the inte- rior of the largest of these islands, particularly in the wilder portions of it, is another race of men with black complexions, crisped hair, and negro faces, called Alfourous. On the coast of New Guinea, and in the neigh- bouring islands, we find other negroes, nearly similar to those of the east- ern coast of Africa, named Papuas*; to the latter, are generally referred the people of Van-Diemen's land, and those of New Holland to the Alfourous. These Malays, and these Papuas are not easily referable to either of the three great races of which we have been speaking ; but, can the former be clearly distinguished from their neighbours, the Caucasian Hindoos and the Mongolian Chinese ? As for us, we confess we cannot discover any sufficient characteristics in them for that purpose. Are the Papuas ne- groes, which may formerly have strayed into the Indian ocean ? We pos- sess neither figures nor descriptions sufficiently precise to enable us to answer this question. The northern inhabitants of both continents, the Samoiedes, the Lap- landers, and the Esquimaux, spring, according to some, from the Mongo- lian race, while others assert that they are mere degenerate offsets from the Scythian and Tartar branch of the Caucasian stock. We have not yet been able to refer the Americans to any of the races of the eastern continent ; still, they have no precise or constant character which can entitle them to be considered as a particular one. Their cop- per-coloured complexion is not sufficient; their generally black hair and scanty beard would induce us to refer them to the Mongoles, if their de- fined features, projecting nose, large and open eye, did not oppose such a theory, and correspond with the features of the European. Their lan- guages are as numberless as their tribes, and no demonstrative analogy has as yet been obtained, either with each other, or with those of the old world "(~. * With respect to the various nations of the Indian and Pacific oceans, see the dissertation of Messrs. Lesson and Garnot in the Zoologie du Voyage de la Coquillc, p. 1- — 113. For the languages of the Asiatics and their affinities, consult the Asia Polyglotta of M. Klaproth. f See the Voyage de M. de Humboldt, and the dissertations of Vater and Mitchill. 4(> MAMMALIA. ORDER II. QUADRUMAXA. Independently of the anatomical details which distinguish it from man, and which have been given, this family differs from our species in a very remarkable way. All the animals belonging to it have the toes of the hind feet free and opposable to the others, and the toes are all as long and flexible as fingers. In consequence of this, the whole species climb trees with the greatest facility, while it is only with pain and difficulty they can stand and walk upright; their foot then resting on its outer edge only, and their narrow pelvis being unfavourable to an equilibrium. They all have intestines very similar to those of man; the eyes directed forwards, the mamma? on the breast, the penis pendent. The brain has three lobes on each side, the posterior of which covers the cerebellum, and the tem- poral fossae are separated from the orbits by a bony partition. In every thing else, however, they gradually lessen in resemblance to him, by as- suming a muzzle more and more elongated, and a tail and a gait more like that of quadrupeds. Notwithstanding this, the freedom of their arms and the complication of their hands allow them all to perform many of the ac- tions of man as well as to imitate his gestures. They have long been divided into two genera, the Monkeys and the Lemurs, which, by the multiplication of secondary forms, have now be- come two small families, between which we must place a third genus, that of the Ouistitis, as it is not conveniently referable to the one or the other. Simia, Lin. The monkeys are all quadrumana, which have four straight incisors in each jaw, and flat nails on all the extremities ; two characters which ap- proximate them more nearly to man than the subsequent genera; their molares have also blunt tubercles like ours, and their food consists chiefly of fruits. Their canine teeth, however, being longer than the rest, sup- ply them with a weapon we do not possess, and which require a hollow in the opposite jaw, to receive them when the mouth is closed. They may be divided, from the number of their molar teeth, into two prin- cipal subgenera, which are again subdivided into numerous groups*. The * Buffon subdivided the monkeys into five tribes: the true monkeys, without tails; the baboons, with short tails; the guenons, with long tails and callous buttocks; the sapajous, with long prehensile tails and no callus; the sagouins, with long tails, not prehensile and without callus. Erxleben, adopting this division, translated these names by simia, pnpio, cercnpithecus, cebus, and callitkrue. Thus it is, that the names QUADRUMANA. 47 Monkeys, properly so called, Or those of the eastern continent, have the same number of grinders as Man, but otherwise differing from each other by characters, which have formed the grounds of the following subdivisions : — The Simia, Erxl. — Pithecus, Geoffr. The Ourangs* are the only monkeys of the ancient continent which have no callus on the buttock ; their hyoid bone, liver, and caecum resem- ble those of Man. Their nose is not prominent, they have no cheek- pouches, nor a vestige of a tail. Some of them have arms long enough to reach the ground when standing — their legs, on the contrary, are very short. S. satyrus, L. ; Audeb., pi. 2; Fr. Cuv. pi. 2. (The Ourang- Outang-j\) Of all animals, this Ourang is considered as approach- ing most nearly to Man in the form of his head, height of forehead, and volume of brain ; but the exaggerated description of some au- thors respecting this resemblance, are partly to be attributed to the cebus and callithrix, by which the antients designated monkeys of Africa and India, have been transferred to those of America. The genus Papio, founded solely on the shortness of the tail, could not be retained, as it violated natural affinities, and all the others required subdividing. It was also necessary to abolish the genus Ouistitis, which was comprised in that of the Sagouins, but which does not altogether corres- pond with the common characters of the other monkeys. * Orang (a) is a Malay word signifying reasonable being, which is applied to man, the ourang-outang, and the elephant. Outang means wild, or of the woods; hence Wild Man of the Woods. f The only good figure of the Ourang- Outang we had for a long time was that of Vosmaer, taken from a living specimen at the Hague. That of Buffon, Suppl. VIII. pi. 1, is every way erroneous; that of Allamand (Buff. d'Holl. XV. pi. 11,) is some- what better — it was copied in Schreber, pi. 2, B. That of Camper, copied ib., pi. 2, C, is tolerably exact, but is easily discovered to have been taken from the dead body. Bontius, Med. Ind. 84, gives a completely ideal one, although Linnaeus took it for the type of his Troglodyte (Amaen. Ac. VI, pi. 1, § 1). There are some good ones in Griffith, and in Krusenstern's Voyage, pi. 94 and 95, but all of them from young subjects. (£=T (a) The species which constitute the sub-genus " Orangs" of Cuvier, are sepa- rated into two sub-genera by Geoffroy, who makes the Simia Satyrus the type of his first sub-genus, Pithecus; and Simia Troglodytes that of his second sub-genus, Troglo- dytes. Besides the distinctions between these two species, described by both Cuvier and Geoffroy, there are two others, which may be easily ascertained on an examina- tion of the skeletons of both. In the Pithecus, or Simia Satyrus, the ribs are of the same number as those of the human body, namely, twelve on each side. But, in the Simia Troglodytes, the ribs on each side are thirteen, the extra pair being arti- culated with the first lumbar vertebra on each side. Between the sternum (breast- bone) of the two apes, a striking difference also prevails. That of the Simia Satyrus is much broader in proportion to its length; and the second, third, fourth, and fifth bones which compose it, are divided longitudinally into two parallel rows, the sepa- rate portions alternating with each other, leaving an indented suture between them, which is peculiarly manifest in the young animal. Now, in the Simia Troglodytes, the sternum is simply divided, in the ordinary way, into five separate portions which are entire; it is altogether much narrower or more compressed laterally than it is in the former species. (See several specimens in the Museum of the College of Sur- geons, in London. — See. also, specimens in the British Museum). — Eng. Ed. 48 MAMMALIA. fact of their being drawn from young individuals only ; and there is every reason to believe, that, with age, their muzzle becomes much more prominent. The body is covered with coarse red hair, the face blueish, and the hinder thumbs very short compared with the toes. His lins are susceptible of a singular elongation, and possess great mobility. His history has been much disfigured by mingling it with that of the other great monkeys, that of the Chimpanse, in particu- lar. After a strict and critical examination, I have ascertained that the Ourang-Outang inhabits the most eastern countries only, such as Malabar, Cochin China, and particularly the great island of Borneo, whence he has been occasionally brought to Europe by the way of Java. When young, and such as he appears to us in his captivity, he is a mild and gentle animal, easily rendered tame and affection- ate, which is enabled by his conformation to imitate many of our ac- tions, but whose intelligence does not appear to be as great as is reported, not much surpassing even that of the Dog. Camper dis- covered, and has well described two membranous sacs in this animal which communicate with the glottis, that produce a hoarseness of his voice — he was mistaken, however, in imagining that the nails are always wanting on his hinder thumbs. There is a monkey in Borneo, hitherto known only by his skeleton, called the Pongo*, which so closely resembles the Ourang-Outang in the proportions of all his parts, and by the arrangement of the fora- mina and sutures of the head, that, notwithstanding the great pro- minence of the muzzle, the smallness of the cranium, and the height of the branches of the lower jaw, we are tempted to consider him an adult — if not of the species of the Ourang-Outang, at least of one very nearly allied to it. The length of the arms, that of the apo- physes of the cervical vertebra?, and the tuberosity of his calcaneum, may enable him to assume the vertical position, and walk upon two feet. He is the largest monkey known, and in size is nearly equal to Man. Mr. J. Harwood, in the Trans. Lin. Soc. XV. p. 471, describes the feet of an ourang, fifteen English inches in length. This an- nounces a very great stature in the animal to which they belonged, and would have led him to the belief that the Pongo is the adult Ourang-Outang, were it not that the skeleton of the Pongo in the College of Surgeons, at London, has one lumbar vertebra more than those of the Ourangs. This, however, is no objection — the same variation is frequently observed in the human subject. The arms of the remaining Ourangs reach only to the knee. They * Audeb. Singes, pi. anat. 2. This name of Pongo, a corruption of Boggo, which is given in Africa to the Chimpanse, or to the Mandrill, was applied by Buffon to a pretended large species of Ourang-Outang — the mere imaginary product of his com- binations. Wurmb, a naturalist of Batavia, has transferred it to this animal, which he was the first to describe, and of which Buffon never had any idea. See Mem. of the Soc. of Batavia, vol. ii. p. 245. The thought, that it might be an adult Ourang, struck me on examining the head of an ordinary Ourang, whose muzzle projected much more than those of the very young specimens hitherto described. T described it in a memoir read before the Acad, des Sciences in 1818. Tilesius and Rudolphi appear also to have had it. See the Mem. of the Acad, of Berlin, 1824, p. 131. QUADRUMANA. -|!) have no forehead, and the cranium retreats from the crest of the eye-brow. The name of Chimpanses might be exclusively applied to them. S. troglodytes, L. (The Chimpanse)* is covered with black or brown hair. Could any reliance be placed on the accounts of tra- vellers, this animal must be equal or superior to man in stature, but no part of it hitherto seen in Europe indicates this extraordinary size. It inhabits Guinea and Congo, lives in troops, constructs huts of leaves and sticks, arms itself with clubs and stones, and thus re- pulses men and elephants ; pursues and abducts, as is said, negro women, &c. Naturalists have generally confounded it with the Ourang-Outang. When domesticated he soon learns to walk, sit, and eat like a man. We now separate the Gibbons from the Ou- rangs. Hilobates, Blig. The Gibbons have the long arms of the true Ourangs, and the low fore- head of the Chimpanse, along with the callous buttocks of the Guenons, differing however from the latter in having no tail or cheek-pouch. They all inhabit the most remote parts of India. S. lar. L.; Buff. XIV. 2; Onko, Fred. Cuv. pi. 5 and 6, (the Black Gibbon) is covered with coarse black hairs, and has a whitish circle round his face. H. agilis, Fred. Cuv. pi. 3 and 4; Petit Gibbon of Buffon, XIV. 3, (the Brown Gibbon) is brown — the circle round the face is of a pale red; the lower part of the back is of the same colour. The young are of a uniform yellowish white— it is very agile, and lives in pairs — its Malay name, Wouwou, is taken from its cry. S. leucisca, Schreber, pi. 3, B, (the Cinereous Gibbon) is covered with a soft and ash-coloured wool. The visage is black — lives among the reeds, and climbs to the tops of the highest branches of the bam- boos, where it balances itself by its long arms. We might separate from the other Gibbons the Siamang. S. syndactila, Raff., Fred. Cuv., pi. 2, (the Siamang) has the second and third toes of the hind foot united by a narrow membrane, the whole length of the first phalanx. It is black — the chin and eyebrows red — lives in numerous troops, which are led by courageous and vigilant chiefs, which, at sunrise and sunset, make the forest ring with the most frightful cries. Their larynx has a membranous sac connected with it. All the ensuing monkeys of the eastern continent have the liver divided * This is the Quojas morou, or the Satyr of Angola, of Tulpius, who gives a bad figure of it, (Obs. Med., p. 271), and the Pygmy, much better represented by Tyson, (Anat. of a Pygmy, pi. 1), copied by Schreber, pi. 1, B. Scotin had given a toler- able drawing of it. copied Amaen. Acad. VI. pi. 1, fig. 3, and Schreber, 1, C. An in- dividual that lived with BufFon, and which is still preserved in the Museum, is repre- sented, though badly, in the Hist. Nat. XIV. 1, where he is called Jocko. The same specimen is much better in Lecat (Traite du Mouv. Muscl. pi. 1, fig. 1), under the name Qidmpese. Audebert gives the same, hut from the stuffed specimen only — he calls it Pongo. VOL. I. E 50 MAMMALIA. into several lobes ; the caecum thick, short, and without any appendage; the hyoid bone has the form of a shield. Cercopithecus, Erxl., partim. The long-tailed monkeys* have a moderately prominent muzzle (of 60°) : cheek-pouches; tail; callosities on the buttocks; the last of the inferior molares with four tubercles like the rest. Numerous species, of every variety of size and colour, abound in Africa, live in troops, and do much damage to the gardens and fields under cultivation. They are easily tamed. Simla rubra, Gm.; Buff. XIV. 30; Fred. Cuv. 24. (The Pa- tras). Red fawn colour above, whitish below , a black band over the eyes, sometimes surmounted w r ith white — from Senegal. Simla cethiops, L. ; Buff. XIV. 32; Fred. Cuv. 25. (The Col- lared Mangabey). A chocolate brown above ; below and the nape of the neck, whitish; on the head a cap or coif of a lively red; eye-lids white. Buffon says it is from Madagascar, and Hasselquist from Senegal; and in fact Sonnerat declares, there are no monkeys in Madagascar. Simla fuliglnosa, Geoff.; Buff. XIV. 32; Fred. Cuv. 25. (The Mangabey). A chocolate brown, uniform above, fawn coloured be- low; eye-lids white. Buffon says it is from Madagascar, and he believes it to be a variety of the preceding. Simla sabcea, Lin. ; Buff. XIV. 37; Fred. Cuv. 19. (The Green Monkey)-]-. It is greenish above, whitish beneath ; face black ; the tufts on the cheeks yellowish ; tip of the tail yellow. From Senegal. Simla j aunus, Gm. ; Malbrouc, Buff. XIV. 29; Simla cynosorus, Scopol. ; Schr. pi. 14, C; Fred. Cuv. pi. 22, var. of the callithrix; Audeb. 4th fam. 2d sect. pi. 5 J. Greenish above; limbs ash-co- loured; face flesh-coloured; no yellow on the tail; one black, and one white band over the eye-brows; scrotum of a beautiful ultra- marine. Simla erythropyga, Fred. Cuv. pi. 21. (The Vervet) differs from the Malbrouc in the scrotum ; which is surrounded with white hairs, the anus with red ones ; and from the Grivet (aS*. grised) Fred. Cuv. 21, by a green scrotum, encircled with fawn-coloured hairs. Simla melarhlna, Fred. Cuv. pi. 18; Buff. XIV. pi. 10. (The Talapoin). Greenish above; tufts of the cheek yellowish; a black nose in the middle of a flesh-coloured face. Sim. mona and S. monacha, Schreb. ; Buff. XIV. 36; Fred. Cuv. 13. (The Mona). Body brown; limbs black; the breast; insides of the arms, and circumference of the head whitish ; black band across the forehead; a white spot at each side of the root of the tail. * Cercopithecus, i. e. tailed monkey, a name used by the Greeks. t Callithrix, Pliny, 1. 8, c. 54, is the name of an Ethiopian monkey, furnished with a beard and a tufted tail, probably the Ottanderou. Buffon arbitrarily applied it to tbis species. X The Cercop. barbalits of Clasius, which Linn, cites as an example of his faunus, is rather an Ouanderbu than a Malbrouc. QUADRUMANA. 51 Sim. diana, Lin; Exquima, Marcgr.*; Audeb. 4th fam. sec. 2, pi. G, and Buff. Supp. VII. 20. (The Roloway). Blackish, speck- led with white above, beneath white ; crupper of a purplish red ; face black, surrounded with white ; a little white beard on the chin. Sim. cephus, Lin.; Buff. XIV. 34; Fred. Cuv. 17. (The Moustache). Ashy-brown ; a yellow tuft before each ear ; a clear blue band, resembling a reversed chevron, on the upper lip. S. petaurista, Gm. ; Audeb. ib. XIV; Fred. Cuv. 13. (The White-nosed Monkey). Black or brown, speckled with white ; white nose ; face black ; circumference of the lips and the eyes reddish. These last five species, all small, beautifully variegated in colour, and of a mild and gentle disposition, are very common in Guinea")". Sejinopithecus, Fred. Cuv. Differs from the Long-tailed Monkeys, by having an additional small tubercle on the last of the inferior molares. They inhabit eastern coun- tries, and their long limbs and very long tail give them a very peculiar appearance. Their muzzle projects very little more than that of the Gib- bons, and, like them, they have callosities on the buttocks. They appear, likewise, to have no cheek-pouches ; their larynx is furnished with a sac. The one longest known is the Sim. nemceus, L.; Buff. XIV. 41 ; Fred. Cuv. pi. 12. Remark- able for its lively and varied colouring ; body and arms grey ; hands, thighs, and feet, black ; legs of a lively red ; the tail and a large tri- angular spot upon the loins, white ; face orange ; he has a black and red collar, and tufts of yellow hairs on the sides of the head ; inhabits Cochin China J. Another species is remarkable for the very extraordinary form of the nose — it is the S. nasica, Schr. ; Buff. Supp. VII. 11 and 12. (The Kahau). Yellow tinted with red; nose extremely long and projecting, in the form of a sloping spatula. This monkey inhabits Borneo, lives in great troops, which assemble morning and evening, on the branches of the great trees on the banks of the rivers— its cry kahau. It is also said to be found in Cochin China. S. entellus, Dufres. ; Fred. Cuv. pi. 8 and 9. (The Entellus). A light yellowish grey ; black hairs on the eye-brows and sides of the head, directed forwards. From Upper Bengal. Is one of the spe- cies held in veneration by the Brahmins. * The figure annexed to the description of the Exquima in Marcgrave is that of an Ouarine, and that of the Exquima is joined to the description of the Ouarine or Guariba. This transposition has produced many errors in synonymes. f Pennant has described certain Guenons without thumbs, Sim. polycomos and Sim. ferruginea, from which Illiger has constructed his genus Colobus, hut I have not yet been able to see them, and for this reason have not mentioned them. M. Tennninck assures us that their head and teeth resemble those of a Semnopithecus. % M. Diard having transmitted to the Museum several Dotics, from Cochin China, it has been proved that they have callosities on the buttocks; a fact denied by Hul- fon, on account of his having seen but one specimen injured by stuffing. The genus Lasiopyga of Illiger must be suppressed, as it is based on this error; E 2 [)% MAMMALIA. S. melalophos, Raff. ; F. C. pi. 7. (The Simpai). Fur of a very lively red ; beneath white ; face blue ; a crest of black hairs reaching from one ear to the other. aS\ comata, Desm. ; S. cristata, Raff. ; Fr. Cuv. pi. 2. Presbitis mitrata, Kotzeb. (The Croo). Fine ash colour below, and the tuft of the tail white ; black crest on the eye-brows, and the hairs of the top of the head long and turned up, forming a tuft. . S. maura, L. ; F. Cuv. pi. 10. (The Negro Monkey). All black, the young of a brownish yellow. The three latter species are from the straits of Sunda*. Macacus-j-. All the animals of this denomination have a fifth tubercle on their last molares, and callosities and cheek-pouches like a Guenon. The limbs are shorter and thicker than in a Semnopithecus ; the muzzle more pro- jecting, and the superciliary ridge more inflated than in either the one or the other. Though docile when young, they become unmanageable when old. They all have a sac which communicates with the larynx under the thyroid cartilage, and which, when they cry out, becomes filled with air. Their tail is pendent, and takes no part in their motions : they produce early, but are not completely adult for four or five years. The period of gestation is seven months — during the rutting season the labia pudendi, &c. of the females are excessively distended J. They are generally brought from India. Sim. silenus and leonina, L. and Gm. ; Ouanderou, Buff. ; Audeb. 2dfam. sect. 1, pi. 3. (The Maned Macaque). Black; ash coloured mane and whitish beard which surround the head. From Ceylon. Sim. sinica, Gm. ; Buff. XIV. 30 ; Fr. Cuv. 30. (The Chinese Monkey). A lively fawn-coloured brown above, white beneath; flesh-coloured face ; the hairs on the top of the head arranged in radii forming a sort of hat. From Bengal, Ceylon. S. radiata, Geoff. ; Fr. Cuv. 29. (The Cape Monkey). Differ- ing from the preceding in a greenish tint. Sim. cynomolgus and cynocephalus, Lin. ; Macaque, Buff. XIV. 20; Fr. Cuv. 26 and 27. (The Hare-lipped Monkey). Greenish above, yellowish or whitish below ; ears and hands black ; face and scrotum tawny §. The Aigrette, Sim. aygula, Lin., Buff. XIV. 21, appears to be a mere variety of this one, differing by a longer tuft of hair on the top of the head. * There is some variation in their Malay names. Raffles, (Linn. Trans. XIII) calls the S. conata, Chinkau; the S. maura, Lotong. Raffles calls the S. fascicularis, the Kra. \ Macaco is the generic appellation of monkeys oh the coast of Guinea, and among the negroes transported to the colonies. Marcgrave mentions a species, which he says has " nares elatas bifidas" — and these vague words, copied from him only, have remained in the character applied to the Macaque of Buff., although it has nothing like it. X Hence the observation of Julian, that monkeys are to be seen in India which have a prolapsus uteri. § Add the Black-faced Macaque, Fr. Cuv. Mammif. 28, and the other species de- scribed in the same work. QUADRUMANA. 5,i Some of the Macaques are distinguished hy a short tail. M. rhesus; Rhesus, Audeb. fam. ii; Patas a queue courte, ib. pi. 4, and Buff. Supp. XIV. pi. 16 ; the first baboon figured by Buff. XIV. pi. 19*. (The Pig-tailed Baboon). Greyish; a fawn-co- loured tinge on the head and crupper, sometimes on the back; face flesh-colour; tail reaching below the hamstrings. From Bengal "f\ Sim. menestrinus, L. ; Sim. platypigos, Schreb. ; Audeb. fam. ii, sect. 1, pi. 2. ; Fr. Cuv. Mammif. under the name of Singe a queue de cochon. (The Brown Baboon). Deep brown above ; black band beginning on the head, and fading as it extends along the back ; yel- lowish round the head and limbs; tail thin and wrinkled J. Inuus, Cuv. Mere Macaques, which have a small tubercle in lieu of a tail. S. silvanus, pithecus and inuus, Lin. ; Buff. XIV. 7, 8 ; Fr. Cuv. Mammif. (The Barbary Ape). Completely covered with a light grey-brown hair, and of all monkeys, is the one that suffers least from our climate. He is originally from Barbary, but is said to have become naturalised in the most inaccessible parts of the rock of Gibraltar §. Cynocephalus, C. II The Dog-headed Monkeys, together with the teeth, cheek-pouches and callosities of the Inuus, Cuv., have an elongated muzzle truncated at the end, in which the nostrils are pierced, giving it a greater resemblance to that of a dog than of any other monkey; their tail varies in length. They are generally large, ferocious and dangerous animals, found mostly in Africa. * The two specimens used by Audebert are still in the Museum. I have exa- mined them and find they are both of one species. f The Macaque a queue courte of Buff". Supp. VII. pi. 13, (Sim. erytrhcca, Schr.) appears to me to be a true Macaque (S. cynomolgus), whose tail had been amputated. X Add the Macaque de I'lnde, and the Macaque a face rouge, Fr. Cuv. Mammif. § The Pitheque of Buff". Supp. VII. pi. 4 and 5, was a young Magot(a). His Lit- tle Cynocephalus, ib. pi. 6, and the Great and Little Cynocephala of Prosper Alpin are also of that species. nSrixo; is the Greek term for monkeys in general, and the one whose anatomy has been given by Galen was a Magot, although Camper thought it was an Ourang-Outang. M. de Blainville perceived this mistake, and I have proved it by comparing with these two species all that Galen has stated respecting the ana- tomy of his pithecus. || Cynocephalus, dog's head, a name well known to the ancients, especially as the dog played a conspicuous part in the symbols of the Egyptians, in which it repre- sented Tot or Mercury. (J5" (a) The Pigmy, or Barbary ape, of which species a male and female are in the Surrey Zoological Gardens, is distinguished in India as an object of superstitious re- verence, to which temples have been raised. In the confined state these animals will- ingly received every sort of food, with the exception of that of animal; they scarcely ever eat any portion, before they broke the whole. The male was capricious, and sometimes ill tempered, and we have seen the female always acting in such a man- ner that shewed fear as well as gentle submission; she usually approached the male by proceeding around him in a circular walk, and with her eyes constantly upon him, as if to watch the favourable moment for shortening the distance between them. The jealousy shewn by him when a visitor took notice of the female, was instantly mani- fested by repeated blows. — Eno. Ed. 51' MAMMALIA. C. papio, Desm. ; Sim. sphynx, Lin. ; Papion, Bull'. (The Gui- nea Baboon). Yellow, verging more or less on a brown ; tufts of the cheeks fawn-coloured; face black; tail long*. They are found of various sizes, owing probably to the difference of age ; when full grown, frightful from their ferocity and brutal lubricity. From Guinea. There is another neighbouring species with a shorter tail, a greener fur, whiter cheek-tufts and a flesh-coloured face, »S'. cynocephalus ; the Babouin, Fr. Cuv. Mem. du Mus. IV. pi. 19. C.porcarius; Sim. porcaria, BoM. ; S. ursina, Penn. ; S.sphyu- giola, Herm. ; The Long-faced Guenon, Penn., and Buff. Supp. VII. pi. 15 ; Black Monkey of Vaillantf ; Chacma, Fr. Cuv. Mammif. Black, with a green or yellowish glaze, particularly on the forehead; tufts of the cheeks grey; face and hands black; his tail reaches his heel, and ends in a tuft of hair. The adult has a large mane — in every thing else, as to habits and form, he resembles the preceding. From the Cape of Good Hope. C. hamadryas ; Tartarin of Belon, Ois. fol. 101, or Papion a perruque; Sim. hamadryas, L. ; Dog-faced Baboon, Penn.; Singe de Moco, Buff. Supp. VII. 10 \. A slightly blueish ash-colour; hairs of the ruff, and particularly those of the sides of the head, very long; face flesh-coloured. This great monkey is also among the most libidinous and horribly ferocious of his kind — lives in Arabia and Ethiopia. There is another species, the Phillippines, which should be dis- tinguished from other Cynocephala, which is totally black, and with- out a tail — S. nigra, Cuv. ; but whose head resembles that of the rest. The Mandrills, Of all the monkeys, have the longest muzzle (30"); their tail is very short ; they are very brutal and ferocious ; nose as in the preceding. Sim. maimon and mormon, Linn. ; Boggo, Choras, Buff. XIV. XVI. XVII. et Supp. VII. 9. (The Mandrill). Greyish brown, inclining to olive above ; a small lemon-yellow-coloured beard on the chin; cheeks blue and furrowed. The nose in the adult male becomes red, particularly at the end, where it is scarlet, which has been the cause of its being deemed, erroneously, a distinct species §. * Those which have been figured as having it short, as the Papions of Buff'. XIV. pi. 13 and 14, &c. had it cut off'. M. Brongnard was the first who gave a good figure of it, but under the improper name of Sim. cynocephalus. His figure is copied by Schreber, pi. 13, B. See the different Papios in the Mammif. Fred. Cuv. f All these factitious species have been established on the good or bad condition of individual specimens of the same species, or on their difference of age. X Copied by Schreber, but badly coloured. There is now a good figure of it in the Mammif. of Fred. Cuv. § We have seen, as well as M. Geoffroy, two or three Mandrills, or S. maimon, change to the Choras or S. mormon, in the menagerie of the Museum. The tuft of hair, which is frequently given as a character of the mormon, is often also in the maimon. QUADRUMANA. 55 The genital parts, and the circumference round the anus, are of the same colour. The buttocks are of a beautiful violet. It is difficult to imagine a more hideous or extraordinary animal. He nearly at- tains the size of a man, and is a terror to the negroes of Guinea. Many details of his history have been mixed up with that of the Chimpanse, and consequently with that of the Ourang-Outang. Sim. leucophcea, Fred. Cuv. Ann. du Mus. d'Hist. Nat. IX. pi. 37, from a young specimen from India, and Hist, des Mammif. from the adult. (The Drill). Yellowish grey; face black; tail very short and thin; in old ones the fur becomes darker, and the chin of a brilliant red. The Monkeys of the New Continent Have four grinders more than the others — thirty-six in all; the tail long ; no cheek-pouches ; buttocks hairy ; no callosities ; nostrils opening on the sides of the nose, and not underneath. All the great Quadrumana of America belong to this division. Their large intestines are less in- flated, and the caecum longer and more slender than in those of the eastern continent. The tails of some of them are prehensile — that is, its extremity can twist round bodies with sufficient force to seize them as with a hand. They are more particularly designated by the name of Sapajous, Cebus, Erxleben*. At their head may be placed the Alouattes (Mycetes, Blig.\ which are distinguished by a pyramidal head, the upper jaw of which descends much below the cranium, as the branches of the lower one ascend very high for the purpose of lodging a bony drum, formed by a vesicular in- flation of the hyoid bone, which communicates with the larynx, and gives to their voice an astonishing volume, and a frightful sound. Hence their name of Howling Monkeys. The prehensile portion of the tail is naked beneath. There are several species, whose distinguishing characters are not yet well ascertained, for the colour of the fur on which they are established varies with the age and the difference of sexes. Simla seniculus, Buff. Supp. VII. 25. (Red Howling Monkey). It is often sent to us from the forests of Guiana, where it lives in troops; size that of a large fox; colour, a reddish chesnut, rather deeper at the head and tail. The Allouatte ourson (Stentor ur sinus, Geoff.), Humb. Obs. Zool. I. pi. 30, must differ from it very slightly ; but it would appear that there are many others, some of which are black or brown, others of a pale colour. In certain species this pale tint is peculiar to the femalesy. * Cebus or Cepus, K^no;, names of an Ethiopian monkey, which, from the de- scription of /Elian, lib. xxvii. c. 8, must have been the Patas. f Marcgrave, Braz. 226, speaks of a black Guariba, with brown hands, that Spix thought he had found in his Seniculus niger. Mem. de Murric, for 1813, p. 333. Mycetes rufimanus, Kuhl. Marcgrave, 227, speaks of another species, all black and bearded, fig. p. 228, un- der the wrong name of Exquima, which must have been, it is probable, the Mycetes 56 MAMMALIA.. The Common Sapajous have the head flat, and the muzzle slightly prominent — facial angle 60°. In some of them, the anterior thumbs are either totally, or nearly so, hidden under the skin, and the prehensile part of the tail naked beneath. M. Geoff, has formed them into a genus by the name of Ateles*. The first species, the Chamek, Ateles pentadactylus, Geoff., differs again from the others in having a slight projection of the thumb, though it is only of one phalanx, but without a nail; its fur is black. A second species, the Mikiri, At. hypoxanthus, Pr. Max.; Brachyteles macrotarsus, Spix, pi. i., has also a very small thumb, and sometimes even a nail. The fur is yellowish, ferruginous to- wards the tail. These two species are separated by Spix under the name Brachyteles. They connect the Ateles with Lagothrix. The other Ateles to which alone Spix restricts that name — Coaita, Buff. — have no apparent thumb whatever. Such are the following: A. paniscus; Simla panisc. L. ; Coaita, Buff. XV. 1. (The Coaita). Completely covered with black hair, like the Chamek, but without any visible thumb ; face, flesh-colour. A. ater, Fr. Cuv. Mammif. (The Cayou). Face black, like the rest of the body. A. marginatus, Geoff. The Chuva, Humb. or the Coaita a face hordee, Ann. Mus. XII. pi. 10. Black, with a border of white hairs round the face. A. belzebuth; Sim. beelzeb., Briss. The Marimonda, Humb. or Coaita a ventre blanc, Geoff.; Ann. Mus. VII. pi. 16. Black above; white beneath; circumference of the eyes flesh-coloured. A. arachnoides, Geoff. Ann. Mus. XIII. pi. 9. (The Spider Monkey). Grey, fawn-coloured or red; eyebrows black. All these animals are natives of Guiana or Brazil ; their fore-feet are very long and slender, and their gait remarkably slowj. bnrbntus, Spix, pi. 32. The female, ib. pi. 33, is of a light yellowish grey. The male must be the Mycetes niger of Kuhl and Prince Maximil. de Neuwied. The Caraia of d'Azzara, which is black, breast and belly of a dark red, the female brownish, may be referred to this species. Pr. Max. has another Mycetes ursinus, which appears to be much browner than the ursinus of M. Geoffroy, and to approximate nearer to the M.fuscus, or the M. discolor of Spix, pi. 30 and 34. This latter rather appears to be the St.fuscus of Geoffroy. The Straw-coloured Alouatte, Stentor stramineus, Geoff, and the Myc. stramineus, Spix, pi. 31, of a yellowish grey, appears from its cranium to be of a different spe- cies, but it may merely be the female of a preceding one. It is easily seen, also, that if their characters are so uncertain, their synonymes must be much more so. Add the St. flavicaudatus, Geoff', of a black brown, with a yellow streak on each side of the tail. * Ann. du Museum, VII. 2fi0, rt seq. f They exhibit some remarkable resemblances to man in their muscles. Of all . animals, they alone have the biceps of the thigh made like ours. QUADRUMANA. [) i Lagothrix(«), Geoff. — Gastrimargus, Spix. Head round, like the Ateles; a thumb developed like the Alouattes; tail partly naked, like the one and the other. Such are the L. Humbol- dii, Geoff. ; the Caparo, Humb. ; Gast. olivaceus, Spix, pi. 28 (the Cap- paro); and the Grison, (or Lag. canus, Geoff.); or Gastr. infumatus, Spix, 29. (The Silver-haired Monkey). Monkeys from the interior of South America, said to be remarkable gluttons. The other Sapajous (Cebus, Geoff.) have a round head, distinct thumbs, and the tail hairy, though prehensile. The species are more numerous than those of the Alouatte, and are characterized with nearly as much difficulty. Some of them have the hairs on the forehead of a uniform length, such as the — Sim. appella, L. (The Sajou); and the S. capucina, L. ; Buff. XV. 4, 5, and 8, 9. (The Capuchin). Both of them of different browns; in the first, the circumference of the face is blackish; in the second it is whitish ; but the shade of colour in all the rest of their bodies varies between a brownish black and a fawn-colour, sometimes even a white. The shoulders and breast are however generally lighter, and the calotte and hands darker*. Others, again, have the hairs of the forehead so disposed as to form a kind of aigrette, such as the Sim. fatuellus, Gm. ; Buff. Supp. VII. 29. (The Horned Sajou). This animal has a tuft of black hairs on each side of the forehead j. The disposition of these monkeys is mild and gentle, their motions quick and light, and they are easily tamed. Their name of Weeping Monkeys is derived from their soft plaintive voice. * The Sajous and the Sais vary so much from a brown to a yellow, that, were there not intermediate varieties, we should be tempted to make many species of them. Such is the case with the Sim. trepida, syrichla, lugubris, flavia, L. and Schreb. as well as some of those distinguished by M. Geoffroy, Ann. du Mus. XIX. Ill and 112. Spix has recently, and in our opinion improperly, multiplied them still more. We would refer to the Sajou {Sim. appella, Lin.) the Cebus robustus, Pr. Max., which appears to us to be an old one of that species. The Ceb. macrocephalus, Spix, pi. 1, does not seem to differ from it, so far as regards the species. We refer to the Sai (S. capucina, Lin.) the Sai a gorge blanche, Buff. (S. hypolencos) ; the Cebus libi- dinosus, Spix, 2; the Ceb. xanthosternus, Pr. Max., or the Ceb. xanthocephalus, Spix, 3; the Ceb. cucullalus, id. 6. We should be more inclined to consider as distinct species, the Sajou a pieds dores, Fred. Cuv., the Sajou brun, id. or Ceb. unicolor, Spix, pi. 4; the Sim. flavia, Schreber, 31, B, from which the Ceb. gracilis, Spix, pi. 5, seems to differ only in the stuffing — but that we require numerous observations, made upon the spot which these ani- mals inhabit, before we can hope to establish their species in any other than an arbi- trary manner. f Here should come the Cebus cirrhifcr, Geoff, and the Ceb. of the same name, of Pr. Max., but which is different. Ceb. cr is talus, Fred. Cuv. g^f(«) The existence of this animal was not known until Humboldt discovered it in South America: — He describes it under the name of Simia Lagothrica. A remark ably fine specimen was presented lately to the Surrey Zoological Gardens, which was brought from Para, on the River Amazon, in South America. Its habitation is now considered to be the northern portion of South America, between the Equator and five degrees of north latitude. — Eng. Ed. 58 MAMMALIA. In the Saimiri the tail is depressed, and almost ceases to be prehen- sile ; the head is very much flattened ; in the interorbitar partition of the skeleton there is a membranous space. There is only one known; the Simla sciurea, Buff. XV. 10. (The Saimiri). Size of a squirrel; of a yellowish grey ; fore-arms, legs, and the four extremities of a yellowish fawn-colour ; end of the nose quite black. Those of the American monkeys, whose tails are not at all prehensile, are called Sakis*. Several of them have the tail long and tufted, whence they have been also termed Fox-tailed Monkeys : their teeth project for- wards more than those of the others. They are the Pithecia of Des- marets and Illiger. Simla pithecia, L. ; Buff. XV. 12; Pithecia inusta, Spix, pi. 10. (The Yarke). Blackish ; circumference of the face whitish. Pith, hirsuta, Spix, pi. 8. (The Grey Sakis). Grey; with yel- lowish hands. Simla satanas, Hofmansegg; Humb. Obs. Zool. L. xxvii. (The Black Saki). All black. Pith, rufiventris, Geoff.; Buff. Supp. VII. 31; Pith, capilla- mentosa, Spix, pi. 11. (The Red-bellied Saki). Brown, with a red belly. Spix distinguishes those species whose tails, although tufted, are shorter than the body, by the name of Brachiurus. His Br. Ouaraki, Sp. pi. 8, has a fawn-coloured body ; head, neck, arms, and feet black. To this should be referred, provided always it is another species, the Sim. melanocephala, Humb. Obs. Zool. p. 29 ; yellow, with a black head. In some, also, the Callithrix, Geoff, or Sagouins, Fr. Cuv. the tail is slender, and the teeth do not project. The Saimiri were associated with them for a long time, but the head of the Sagouins is higher, and their canine teeth much shorter. Such are the Call, personata, Geoff., Spix, pi. 12; Call, nigrifrons, id. 15. (The Masked Monkey). A yellowish grey; head and hands black. Call, lugens; S. lugens, Humb. (The Mourning or Widow Mon- key). Blackish, with a large white gorget or neck-piece. The Call, amicta, Geoff., Sp. pi. 13, and the Call, torquata, Hofmansegg, can differ but little from this species j. Nocthorus, Fred. Cuv. — Nyctipithecus, Spix. Improperly called Aotus by Illiger. Only differs from the Sagouins in its great nocturnal eyes, and in the * All the American monkeys, whose tails are not prehensile, together with the Ouistitis, are termed hy Buffon Sagouins (Callithrix, Erxl.) This name of sagouin or cagui is in fact applied in Brazil to all the little Quadrumana, whose tails are not prehensile. N.B.— M. Geoff., Ann. Mus. XIX. 112, 113, gives to his Callithrix, which are merely a division of those of Erxleben — Nocthorus and Pithecia, the common name of Geopithecus. f Add Call.melatwchir, Pr. Max. — C. cinerascens, Spix, pi. 14, is the young of the same according to Temminck. — C. cuprea, Spix, pi. 17. — C. g'go, id. pi. lfi. N B.— This name of Gigo or Guigo is given by Pr. Max. to his Melanochir, so that we must consider it generic. QUADRUMANA. 59 ears, which are partly hidden under the hair. One species only is known. Nocth. trivirgata, Fred. Cuv., Mammif. ; Nyctipith. vociferans, Spix, pi. 18. (The Douroucouli). Ash-coloured above, fawn-co- loured beneath; a black vertical line on the middle of the forehead, and one on each temple. It is a nocturnal animal of South Ame- rica*. They are all from Guiana or Brazil. Ouistitis (a). — Hapale, Illig. — Arctopithecus, Geoff. A small genus, similar to the Sakis, and for a long time confounded with them in the great genus of monkeys. In fact, like the generality of the American monkeys, they have the head round ; face flat ; nostrils lateral ; buttocks hairy; no cheek-pouches, and, like the Sakis in particular, the tail not prehensile. They have only, however, twenty grinders, like the monkeys of the ancient continent; all their nails are compressed and pointed, those of the hind thumbs excepted, while their anterior ones are so slightly separated from the fingers, that it is with hesitation we assign to them the name of quadrumana. They are all pretty little creatures, of agreeable forms, and easily tamed. M. Geoffroy distinguishes the Ouistitis, properly so called, which he names Jacchus, and whose peculiar characters are pointed inferior in- cisors, arranged on a curved line, equal to the canines. Their tail is an- nulated and well covered with hairs ; the ears generally ornamented with a hairy brush. Sim. jacchus, Lin. ; in Paraguay the Titi, Buff. XV. pi. 24. (The Common Ouistiti). Tail tolerably well tufted, coloured by rings of brown and white ; body greyish-brown ; two large tufts of white hairs before the ears. From nearly every part of South America-f\ M. Geoffroy calls those species which have inferior trenchant incisors placed nearly in a straight line, and less than the canines, Midas. Their tail is also more slender and not annulated. Sim. cedipus, L. ; Buff. XV. 17. (The Pinche). Grey, waved with brown; long white hairs on the head which hang behind the ears; tail slender and red. From the banks of the Amazon t. * Add Nyctipithec.felinus, Spix, pi. 18. f It is difficult to establish very specific limits between Ouistitis of different co- lours. The Jacch. penicillatus, Geoff., Spix, pi. 26, has a white spot on the forehead, and the tufts of the ears brown or black. — His J. leucocephalus, Pr. Max., lib. 2, has the same tufts, but the whole head and fore part of the neck are white. — His J. hu- meralijer has the breast, shoulders and arms white. — The J. albicollis, Spix, pi. 25, has the spot on the forehead, tufts of the ears and a large collar all white. In some of them, on the contrary, all the white has disappeared. See Anna! du Mus., XIX. p. 119—122. f I suspect the Mid. bicolor, Spix, pi. 24, is merely a variety of the Sim. cedipus, and his M. my s tax of the M. labiatus. f!3F (a) The name of Ouistitis is given to the animals of this species from the peculiar sound which they emit, and which is very closely imitated if we express separately and at intervals the successive syllables which compose the word. — Eng, Ed. (50 MAMMALIA. Mid. rufimanus, Geoff'.; Sim. midas, L. ; Buff. XV. 13. (The Tamarin). Black, tlie four hands yellowish. From Guiana. Mid. ursulus, Geoff. ; Buff. Supp. VIII. 32 ; Mid. fuscicollis, Spix, pi. 20. (The Black Tamarin). All black ; reddish wavings on the back. Mid. labiatus, Geoff.; M. nigricollis, Spix, 21. (The White- lipped Tamarin). Black; crupper reddish; circumference of the muzzle white*. Sim. rosalia, L. ; Buff. XIV. 16. (Lion Monkey, or the Mari- kina). Yellowish ; the head surrounded with a golden gilt yellow mane ; end of the tail brown. From Surinam. Hapale chrysomelas, Pr. Max. lib. ii. (Black Marikina). Black ; fore-arms and upper side of the tail and mane round the head of a strong golden yellow. Sim. argentata, L.; Buff. XV. 18. (The Mico). Silver grey, sometimes all white ; tail brown. From the Amazon. Lemur («), Lin. The Lemurs, according to Linnaeus, comprehend all the Quadrumana which have in either jaw incisors differing in number from four, or at least differently directed from those of the Monkeys. This negative character could not fail to embrace very different beings, while it did not even unite those which should be combined. Geoffroy has established several divisions in this genus which are much better characterized. The four thumbs of these animals are well developed and opposable, and the first hind finger is armed with a pointed, raised nail; all the other nails are flat. Their fur is woolly ; and their teeth begin to exhibit sharp tubercles catching in each other as in the Insectivora. Lemur.- — Maris, properly so called. Six incisors in the lower jaw compressed and slanting forwards, four in the upper that are straight, the intermediate ones being separated from each other ; trenchant canines ; six molares on each side above ; six be- low ; ears small. They are very active animals, which, from their pointed * The S. leonina, Humb. Obs. I. pi. 5, is brown, with white lips and black face, like this species; but it appears the hairs of the neck are more thickly set, forming a mane like that of the Marikina. Add Mid. cltrysopygus, Natterer. 55T (a) Lemures was the word employed by the Romans to express ghosts which walked by night, and because the animals now called Lemurs were remarkable for their disposition to sleep during the greater portion of the day, whilst at night they always became restless and bounded about with the greatest agility, Linnaeus gave to them the above designation. Besides the peculiar characters of the Lemurs mentioned in the text, there are others which may be seen in the specimens in the London Zoological Gardens, namely, the elongated face, the round and prominent eyes, the long curved nail on the index finger of the hinder hand; they possess scarcely any external character in common with the monkey, save in the prehensile power of the hands. There is reason to believe that the Lemurs are occasionally, or partially, carnivorous, and the nature of their teeth fully justifies the opinion. — Eng. Ed. QUADRUMANA. Gl heads, have been called Fox-nosed Monkeys. Their food is fruit. Their species are very numerous, and are only met with in the island of Mada- gascar, where they appear to replace the monkeys, none of which, it is said, are to be found there. Nearly all the difference that exists between them is in the colour. L. catta, L.; Le Mococo, Buff. XIII. 22. Ashy-grey; tail black, and white rings. L. macaco, L. .■ Le Vari, Buff. XIII. 27. Variegated with large black and white spots. L. ruber, Peron ; Le Mahi rouge, Fr. Cuv. Mammif. A lively reddish chestnut ; head, four hands, tail, and belly black ; a white spot on the nape of the neck, a red tuft to each ear. L. mongos, L. ; Le Mongous, Buff. XIII. 26. All brown; face and hands black; and other neighbouring species or varieties, such as L. albifrons, Geoff. ; Le Mongous a front blanc, Audeb., Makis, pi. 3. Brown; forehead white, &c* Indris. — Lichanotus, IUig. Teeth like the preceding, except that there are only four in the lower jaw. One species only is known ; it has no tail ; is three feet high ; black ; face grey ; posteriors white, (Lemur Indri), Sonnerat. Se- cond Voy., pi. 86. The inhabitants of Madagascar tame and train it like a hound f. Loris. — Stenops, IUig. The Sloth Monkeys have the teeth of the Makis, except that they have sharper points to the grinders ; the short muzzle of a mastiff; body slen- der; no tail; eyes large and approximated; tongue rough. They feed on insects, occasionally on small birds or quadrupeds; their gait is excessively slow, and mode of life nocturnal. M. Carlisle has found at the trunk of the arteries of the limbs the same state of ramifica- tion as is found in the true Sloths. Two species only are known, both of them from the East Indies. Lem. tardigradus, L. (The Loris Sloth, or Sloth of Bengal). Buff. Supp. VII. 36. Fawn-coloured grey, a brown streak along the back; two of the upper incisors sometimes wanting J. * Add the Black Maki, L.; Niger, Edw. 218.— The Black-fronted Maki (/,. nigri- frons, Geoff.) — The Black headed Makis {L. melanocephalus, Fr. Cuv.) — The Straiv- berry Maki, — The Red Maki, Audeb. pi. 2, &c. But it is not certain that many of these species do not resolve themselves the one into the other. See Geoff., Ann. Mus. XIX. p. 160. f The Long-tailed Indri, {Lemur laniger, Gem.) Sonnerat, Second Voy., pi. 87, needs revision. + The slowness of its gait, which caused it to be mistaken for a Sloth, has in- duced some authors to maintain, in opposition to Buffo n and to truth, that the genus of the Sloths exists also in Asia. G2 MAMMALIA. Lem. gracilis(a), L. (The Slender Loris). Buff. XIII. 30, and better, Seb. I. 47. Fawn-coloured grey; no dorsal stripe; a little smaller than the preceding; nose more raised by a projection of the intermaxillaries *. Galago, Geoff. — Otolicnus, Illig. Have the teeth and live on the insectivorous food of the preceding; elongated tarsi which produce a disproportion in the dimensions of their hind feet; a long tufted tail; large membranous ears and great eyes, which imply a nocturnal life. There are several species known, all from Africa : \. It would ap- pear that we should refer to these also an animal of that country (Lemur potto, Gm.), the Bosnian, Voy. in Guin., p. 252, No. 4, whose gait is said to be as slow as that of the Loris and Sloths. Tarsius. The Tarsiers have the tarsi elongated, and all the other peculiarities of form belonging to the preceding division; but the space between their grinders and incisors is occupied by several shorter teeth ; the middle su- perior incisors are lengthened and resemble the canine. The muzzle is very short, and the eyes still larger than those of all the preceding. They are nocturnal animals, and feed on insects. From the Moluccas. Lemur spectrum, Pall., Buff. XIII. 9+. * From this difference in the nose, M. Geoffroy makes of the first species the genus Nycticebus, and of the second that of Loris. f The great Galago, as large as a rabbit (Galago crassicaudatus, Geoff.) The middling one the size of a rat (Galago senegalensis, id.); Schreb. XXXVIII. Bb. Audeb. Gal. pi. 1. — The small one a little less, Brown, 111. 44. — Compare also the Galago of Demidorf, Fischer, Mem. des Nat. de Moscou, I. pi. 1. % Compare the Tarsius fuscomanus, Fischer, Annat. des Makis, pi. 3, and the Tarsius bancanus, Horsfield, Java. Travellers should search for certain animals drawn by Commerson, and which M. Geoffroy has had engraved, Ann. Mus. XIX. 10, under the name of Cheirogaleus. These figures seem to announce a new genus or subgenus of the Quadrumana. (j3T ( a ) In the examination of a specimen of this species which recently died in the Zoological Gardens in Regent's Park, the distribution of the arteries to the limbs was found to be analogous to that very peculiar one which obtains in the Lemurs and Sloths, except, that in the Loris the structure and distribution of the vessels supply- ing the blood appeared to be destined more to the object of increasing the tenacity of the animal's grasp, and to allow to him to prolong the state of muscular contraction with impunity. — Eng. Ed. CARNARIA. Go ORDER III. CARNARIA (a).— CARNIVOROUS ANIMALS. This order consists of a considerable and varied assemblage of ungui- culated quadrupeds, possessing, like Man and the Quadrumana, the three sorts of teeth, but which have no opposable thumb to their fore-feet. Their food is animal, and the more exclusively so, as their grinders are the more trenchant. Those which have them, either wholly or in part tuberculous, live more or less on vegetable substances, and those in which they appear with conical points, live principally on insects. The articu- lation of their lower jaw, having a cross-wise direction, and its parts being combined on the principle of the hinge, is incapable of horizontal motion, and possesses merely the faculty of opening and of closing. Their brain has the usual depressions, but it has no third lobe, nor does it lie upon the cerebellum in these animals any more than it does in the families hereafter to be described; their orbit is not separated from the temporal fossa in the skeleton, the cranium is narrowed and the zygo- matic arches widened and raised, in order to give more strength and vo- lume to the muscles of their jaws. Their predominant sense is that of smell, and their pituitary membrane is generally spread over numerous bony laminae. The fore-arm has the power of rotating in nearly all of them, although with less facility than in the Quadrumana, and they never have in the fore-feet thumbs opposable to the other toes. Their intes- tines are less in volume in consequence of the substantial nature of their food, and in order to prevent the putrefaction which flesh would necessa- rily experience in being kept too long in a canal of great length. Besides, their forms and minute portions of their organization vary considerably, and are the source of analogous varieties in their habits, and to a degree which makes it impossible to arrange their genera upon one common scale, so that it becomes indispensable to form them into several families, which are variously connected together by multiplied relations. ftCf (a) From some experiments recently performed in the Zoological Gardens in Regent's Park, it would appear that Carnivorous Mammalia fed with two meals a day, are by no means in such good condition as those which have the same quantity of flesh in one meal only. Two Leopards were chosen for the first experiment. One, which weighed 911bs., was fed in the usual manner, on 41bs. of beef every day in one meal — the other, which weighed 100^ lbs., was supplied with the same quan- tity of beef, but one-half was given in the morning and the other half in the even- ing. After an interval of five weeks, during which the animals were fed in this way, they were weighed; when it was found that, whilst the Leopard that had his food all at once, gained lib. in weight; the other lost £lb., and his temper became very much worse. Two Hyaenas were subjected to a similar experiment, which was at- tended with pretty nearly the same results. — Eng. Ed. 64 MAMMALIA. THE FIRST FAMILY OF CARNARIA. THE CHEIROPTERA Retain some affinities with the Quadrumana by the pendulous penis, and by the mammae which are placed on the chest. Their distinguishing- character consists in a fold of skin, commencing at the sides of the neck, and extending between the four members and ringers of the anterior feet, supports them in the air, and even enables such of them to fly as have their hands sufficiently developed for that purpose. This disposition required strong clavicles and large scapula? to give the necessary solidity to the shoulder, but it was incompatible with the rotation of the fore-arm, which would have diminished the force of the effort requisite for flight. They have all four great canines, but the number of their incisors varies. They have long been divided into two genera, founded upon the extent of their organs of flight; but the first of these requires several subdivisions. Vespertilio, Lin. The Bats have the arms, fore-arms, and fingers excessively long, form- ing, with the membrane that occupies their intervals, true wings, possess- ing even a greater extent of surface than those of Birds — they conse- quently fly very high, and with great rapidity. The thickness of their pectoral muscles is proportioned to the motions they have to execute, and there is a ridge in the middle of the sternum like that of Birds, to which they are attached. The thumb is short and armed with a claw, by which they are enabled to creep and to suspend themselves. Their hind feet are weak and are divided into five toes, almost always of equal length, armed with trenchant and pointed nails. They have no caecum in their intes- tines. Their eyes are excessively small, but their ears are frequently very large, and together with the wings form an enormous membranous surface, which is almost naked, and so extremely sensible, that the Bats are conducted through all the sinuosities of their labyrinths, even after their eyes have been plucked out, probably by the diversity of the im- pressions of the air. They are nocturnal, and in our climate pass the winter in a state of lethargy. During the day they suspend themselves in obscure places. They generally produce two young ones at a birth, which they keep fastened to their mammae, and whose size is consider- able in proportion to that of the mother. This genus is very numerous, and offers many subdivisions. We must begin by separating from it the Pteropus, Briss. The Bats called the Roussettes, have trenchant incisors in each jaw, and grinders with flat crowns*; their food, consequently, consists chiefly * The grinders have properly two longitudinal and parallel projections separated by a groove, wliich wear away by attrition. CARTS ARIA. 65 of fruit, of which they destroy considerable quantities; they know, how- ever, how to pursue birds and small quadrupeds. They are the largest Bats known, and their flesh is eaten. They inhabit the East Indies. Their membrane is deeply notched between their legs ; they have no tail, or nearly none ; the index finger, which is but half the length of the medius, has a third phalanx, and a little nail which is wanting in the other Bats; each of the other fingers, however, has but two phalanges. The muzzle is simple, the nostrils are widely separated, the ears are of a middling size, but without a tragus (a), and the tongue is bristled with points that curve backwards ; the stomach is an elongated sac, unequally inflated. They have never been found, except in Southern Asia or the Indian Archipelago. I. The Roussettes without tails, with four incisors in each jaw*. P. edulis, Geoff. (The Black Roussette, or Edible Bat). Blackish brown, deepest beneath, nearly four feet between the extremities of the wings. From the Straits of Sunda and the Moluccas, where they are found in great numbers during the day suspended to the trees. To preserve fruit from their attacks, it is necessary to cover it with nets. Their cry is loud and resembles that of the goose. The Bat is taken by holding to him a bag fastened to the end of a rod; the flesh is esteemed a delicacy by the natives, but Europeans dislike it on account of its musky scent -f. Pter. vulgaris, Geoff.; Buff. X. 14. (The Common Roussette). Brown, face and sides of the back fawn-coloured. From the Isle of France and Bourbon, where it is found on the trees in the forests. Its flesh has been compared to that of the hare and partridge. Pter. rubicollis, Geoff.; Buff. X. 17. (The Red-collared Rous- sette, the Roussette of Buffon). Greyish brown, the neck red. From the same islands, where it lives in the hollows of trees and in holes in rocks J. II. With a small tail and four incisors in each jaw. M. Geoffroy was the first who described the species of this sub- division. One of them woolly and grey, Pter. cegypticus, is found in the caves of Egypt. Another is reddish, and has a somewhat longer tail, half involved by the membrane — Pter. amplexicaudus, Ann. du Mus. torn. XV. pi. 4. From the Indian Archipelago, &c. §. * Linnasus confounded them under his species Vespertilio vampirus. f According to Zemminck, the Roussette of Edwarde Geoff., Edw. 108; it is fawn-coloured, and deep hrown in the back: it is only the young state of this species. % Add Pter. medius; — Pter. phceops ; — Pter. poliocephalus; — Pter. dasymallus ; Temm., Mamm., pi. 10. — Pter. pallidas; — Pter. Keraudrenius, Quoy and Gaym., Voy. de Freycinet; — Pter. griseus, Geoff. Ann. Mus. pi. 3, XV. vi, cop. Temm., pi. 11: — Pter. personatus ; — Pter. melanocephalus, Temm. pi. 12. § Add Pter. stramineiis ; — Pter. marginatus, Geoff, loc. cit. pi. 5; — Pter. minimus, id. or the Kiodote, Fr. Cuv., or the Pter. rostratus, Horsf. (£J* (a) The Tragus is the small prominence of a triangular form, which, in the external portion of the ear, projects over the anterior and outer part of the auditory canal; it forms, in the human ear, the terminating portion of what is called the antihelix. — Eng. Ed. G6 MAMMALIA. III. According to the characters of M. Geoffroy, we further separate from the Roussettes the Cephalotes which have the same kind of grinders, but whose index finger, short, and consisting of three phalanges, like that of the preceding, wants, however, the nail. The membranes of their wings, instead of meeting at the flank, are joined to each other on the middle of the back, to which they adhere by a vertical and longi- tudinal partition. Very often they have hut two incisors. C. Peronii, Geoff. ; Ann. du Mils. XV. pi. 4. (The Cephalote of Peron). Brown or red. From Timor. The Roussettes being withdrawn, we have the true Bats left, which are all insectivorous, and have three grinders on each side in each jaw, bristled with conical points, that are preceded by a variable number of false molars. Their index ringer never has a nail, and, one subgenus ex- cepted, the membrane is always extended between the two legs. They should be divided into two principal tribes. The first has three ossified phalanges in the middle finger of the wing, but the remainder, in- cluding the index itself, consists of but two. To this tribe, which is ahnost exclusively foreign, belong the following subgenera. Molossus, Geoff. — Dysopes, Ill'uj. Has the muzzle simple ; ears large and short, arising near the angle of the lips, and uniting with each other on the muzzle; the tragus short, and not enveloped by the conch (a). The tail occupies the whole length of their inter-femoral membrane, and, more frequently, even extends beyond it. They have almost always two incisors in each jaw, though, according to M. Temminck, several of them have at first six below, of which four are successively lost. The Dinops of M. Savi belong to the Molossus with six inferior in- cisors. There is one species in Italy — Dinops cestonii, Savi, Giorn. de Letter., No. 21, p. 230. M. Geoffroy calls those in which he has counted four inferior incisors Nyctinomus*. The Molossi, at first, were only found in America j-; at present, how- * The Niictinome of Egypt, Geoff, Eg. Mammif., pi. 11, f. 2, and Temm., Monog. des Mammif. pi. 19; — the Nyctinome of Brazil, Isid. Geoff, Ann. des Sc. Nat., I. pi. 22, or Mol. nasutus, Spix, pi. 35, f. 7; — the N. slender tenuis, (Horsfield, Java, N. No. 5), and Temm. Monog. pi. 19, bis. •f Buffon has three of them confounded by Gmel., under the common name of Vespertilio molossus; M. longicaudatus, Buff X. xix, 2; — M. fusciventer, lb. 1; — M. guyanensis, Id. Supp. VII. lxxv. Since then they have been increased. M. rufus, Geoff, Ann. Mus. VI. 155; — M.alecto, Temm., Monog., pi. xx; — M.abrasus, Temm., lb., pi. xxi; — M. velox, Natterer, Temm., pi. xxii, 1; — M. obscurus, Geoff, Temm., lb., pi. xxii, 2. These species, however, have not been sufficiently compared with those of Buffon, nor with the M. ur sinus, Spix, pi. xxxv, f. 4, and the M. fumarius, lb., f. 5 and 6. g_T (a) The conch is a deep conical cavity, situated within the eminences of the outer part of the ear; it is bounded above by a prominent curved margin, which is called the antihelix ; and the conch leads to the canal, through which the sound passes into the interior of the ear. — Eng. Ed. CARNAR1A. (J7 ever, we know several of both continents*. Some of them have the thumb of the hinder feet placed at a greater distance from the first finger than the fingers are from each other, and endowed with a power of sepa- rate motion, a character on which, in a species where it is very strongly marked, M. Horsfield has established his genus Cheiromeles-J"-. It is here, perhaps, that we should also place the Thiroptera of Spix, which appear to have several characters of the Molossi, and whose thumb has a little concave palette peculiar to them, and by which they are ena- bled to cling more closely j. Noctilio, Lin., Ed. XII. Has the muzzle short, inflated, and cleft, as in a double hair-lip, fur- nished with warty tubercles and odd looking seams ; ears separate ; four incisors above, and two below ; tail short, and free above the inter-femoral membrane. The species best known is from America. It is of a uniform fawn-colour — Vespert. leporinus, Gm. Schreb. LX. §. Phyllostoma, Cuv. and Geoff. In which the regular number of incisors is four to each jaw, but in which a part of the lower ones frequently fall, being forced out by the growth of the canines: they are moreover distinguished by a membrane in the form of a leaf, which is reflected crosswise on the end of the nose. The tragus of the ear resembles a small leaf, more or less indented. The tongue, which is very extensible, terminates in papilla?, which appear to be so arranged as to form an organ of suction — the lips also are furnished with tubercles, symmetrically arranged. They are all from America, run along the ground with more facility than the other bats, and have a habit of sucking the blood of animals. 1. The Phyllostomes without a tail. — Vampirus, Spix. P. spectrum; V. spectrum, Lin.; the Andira-guacu of the Bra- zilians; Seb. LVIII; Geoff. Ann. Mus. XV. xii, 4. (The Vam- pire). The nasal leaf wrought into a funnel ; colour a reddish brown ; size, that of a magpie. From South America. It has been accused of causing the death of men and animals by sucking their blood ; but it does no more than inflict very small wounds which may sometimes be affected by the poisonous influence of the climate ||. * M.plicatus; Vespert. plicatus, Buchan. ; Lin. Trans., V. pi. xiii; — Dysopes rup- pelii, Temm., Monog., pi. xviii. f Cheiromeles torquatus, Horsf., Jav. or Dysopes cheiropus, Temm., Monog., pi. xvii. X Thir. tricolor, Spix, 36, f. 9. It is with some hesitation that we have thus placed this subgenus, its description being incomplete. § The N. dorsatus, Geoff., or the N. vittatus, Pr. Max., has a white stripe down the back. — The N.albiventer, Spix, 35, 2 and 4, is fawn-coloured above, white beneath, and rather smaller. Add N. rufus, Spix, 35, 1. || Add the Lunette; Vesp. perspicillatus, L.; Buffi, Supp. VII. lxxiv; and the three species from D'Azara, by Geoff., Ann. du Mus. VI. 181 — 182. 08 MAMMALIA. 2. The Phyllostomes with a tail involved in the inter-femoral membrane. V. hastafus, L. Buff. XIII. xxxiii. (The Javelin Bat). Nasal leaf in the form of a javelin, with entire borders*. 3. The Phyllostomes with the tail free above the membrane. P. crenulatum, Geoff. Ann. du Mus. XV. pi. 10. (The Crenu- lated or Indented Javelin Bat). The nasal leaf in the form of a javelin notched in its border. M. Geoffroy, Mem. du Mus. IV. p. 418, separates from the Phyllos- tomes those species whose tongues are narrow, and extensible, and fur- nished with papillae resembling hairs — he calls them Glossophaga. All species are likewise from America \. In the second great tribe of Bats the index has only one bony phalanx, while all the other fingers have two each. This tribe is also divided into several subgenera. Megaderma, Geoff. Ann. du Mus. XV. Which have on the nose a leaf, more complicated than that of the Phyl- lostomes; the tragus large and most commonly bifurcated; the conchs of the ear very ample and united one with the other on the top of the head ; tongue and lips smooth ; the inter-femoral membrane entire, and no tail. They have four incisors below, but there are none above, and their inter- maxillary bone remains cartilaginous. They are all from the old continent, either from Africa, as the Leaf from Senegal for instance, (Meg. Frons., Geoff), with the nasal leaf oval and nearly as large as the head ; or from the Indian Archipelago, as the Spasma of Ternate, Vespert. Spasma, L., Seb. I. lvi. — La Lyre, Geoff. Ann. Mus. XV. pi. 12. — Le Trifle de Java, Id. ib., &c. They are distinguished from each other by the figure of their leaves, as in the Phyllostomes. Rhinolophus, {Geoff, and Cuv.) commonly called The Horse- Shoe Bats. Which have the nose furnished with very complex membranes and crests laid upon the chanfrin, presenting the figure of a horse-shoe ; the tail is long and placed in the inter-femoral membrane. There are four incisors below, and two very small ones above in a cartilaginous inter-maxillary bone. There are two species of them in France which are very common, discovered by Daubenton. R. bifer, Geoff., Ann. Mus. XX. pi. 5 ; Vesp. ferrum equinum, L., (the Great Horse-Shoe Bat); and Vesp. hipposideros, Bechst. (the Small Horse-Shoe Bat); Buff. VIII. xvn, 2 and 20; Geoff, loc. cit., both of which inhabit quarries, remaining isolated there, * Add Pkilost. elongatum, Geoff., Ann. Mus. XV. ix. f Vespertilio soricinus, Pall. Spicil. Fascic. III. pi. 3 and 4, copied Buff. Supp. III. pi. 53. — Gloss&ph. ample xicaudatus, Geoff. Mem. Mus. IV. pi. 18, F. C. — Gl. caudifer, Id. ib. pi. 17, fig;. A, and B. CARNARIA. 69 suspended by their feet, and enveloping themselves with their wings, so that no other part of the body is visible*. Nycteris, Cuv. and Geoff. The forehead furrowed by a longitudinal groove, which is even marked upon the cranium, bordered by a fold of the skin which partially covers it; nostrils simple ; four incisors without intervals above and six below ; ears large and separated; tail involved in the inter-femoral membrane. They are African species. Daubenton has described one by the name of the Campagnol volant, Buff. X. pi. xx, fig. 1 and 2, the V. hispidus, Lin., Schreb. LVI. M. Geoffroy has found others in Egypt f. Rhinopoma, Geoff. The pit on the forehead less strongly marked; nostrils at the end of the muzzle, and a little lamina above, somewhat resembling a currier's knife; ears united; tail extending far beyond the membrane. One is known in Egypt, where it is principally found in the pyramids J. Taphozous, Geoff. A small round pit on the forehead, but no recurved leaf to the nostrils ; head pyramidal ; only two incisors above, and very often none ; four trilo- bate incisors below; ears wide apart, and the tail free above the mem- brane. The males have a transverse cavity under the throat. A little prolongation of the membrane of the wings forms a sort of sac near the carpus § (a). One species was discovered in the catacombs of Egypt, by M. Geoffroy ||. Mormoops, Leach. Four incisors in each jaw, the superior tolerably large, the inferior tri- lobate ; cranium singularly raised like a pyramid above the muzzle ; on each side of the nose is a triangular leaf which extends to the ear**. Vespertilio, Cuv. and Geoff. The common Bats, or Vespertilions, have the muzzle without leaf or other distinguishing marks ; ears separate ; four incisors above, of which the two middle ones are apart, and six trenchant incisors slightly denticu- * Add the other four species figured Geoff. Ann. Mus. XX. pi. 5, of which one is the Vesp. speoris, Schreb. LIX. B, and Peron, Voy. aux Terres, Aust. pi. 35. f The Thebaic Nyctertis, 29, Mammif. I, 2, 2; and Ann. Mus. XX. pi. 1.— The Javanese Nyctertis, Geoff. Ann. Mus. XX. pi. 1. X Rhinopome microphylle, Geoff.; Vespertilio microphyllus, Schr. § It was this that caused Illiger to name the genus which contained the Taphiens Saccopterix. || The Taphien filet, Eg. Mammif. I. 1, 1.— The Taphien per/ore, lb. III. l, which does not appear to differ from the Flying Lerot, Lerot Volant, Daub.; — T. senega- lensis, G. — Add the Vesp. lepturus, Gm., Schr. LVI I. — The T. of India; V. brach- manus, G. — The T. of the Isle of France; T. mauritianus, G. — The T. rnfus, Wils. Amer. Ornith. vol. VI. pi. 50, No. 4. — The T. lonsimamis, Hardw. Lin. Trans, vol. and pi. XVII. ** The species — Mormoops Blainvillii, Leach, Lin. Trans. XIII — is from Java. (a) The carpus is the hand. — Eng. Ed. 70 MAMMALIA. lated below ; the tail involved in the membrane. This subgenus is the most numerous of the whole, its species being found in every part of the world. France alone has six or seven. The tragus of some is shaped like an awl, and to this division belongs the most commonly known species. Vesp. murinus, L. ; V. myotis, Kuhl, Buff. VIII. xvi. (The Common Bat). Oblong ears, the length of the head; hair brown; maronne above, bright grey beneath ; the young of an ashy grey. Some other smaller but neighbouring species have lately been ob- served in Europe*. In others again the tragus is angular, such as the Vesp. serotinus, L. ; Buff. VIII. xviii, 2. (The Serotine Bat). A deep maronne ; wings and ears blackish ; the conch triangular and shorter than the head. The female is paler than the male. Found under the roofs of churches, and of other little frequented edifices, &c.f- A third kind has a crescent shaped tragus. V. noctula, L. ; Buff. VIII. xviii, 1; V. proterus, Kuhl; V. la- siopterus, Schreb., 58, B. (The Noctule Bat). Fawn coloured; ears triangular, shorter than the head ; tragus rounded, a little larger than the preceding. Found in the hollows of old trees, &c. V. pipistrellus, Gm. ; Buff. VIII. xix, 1. (The Pipistrille). The smallest one in France ; a blackish brown ; ears triangular J. M. Geoffroy separates still further from the Vespertilio, the Plecotus, Geoff. Oreillards, whose ears are larger than the head, and are united to each other on the cranium, as in Megaderma, the Rhinopomes, &c. ; the tragus large and lanceolate — and there is an operculum on their auditory aper- ture. The common species — Vesp. auritus, L.; Buff. VIII. xvn, 1. (The Long-eared Bat). Still more abundant in France than the Bat. Its ears are nearly as large as the rest of the body. It lives in houses, kitchens, &c. There is also another discovered by Dau- benton — (the Barbastelle) — Vesp. barbastellus, Gm., Buff> VIII. 19, 2. Brown, with much smaller ears§. * The V. Bechsteinii, Leisler, Chauves. d'Allem., pi. 22. — The V. mijstacinus, lb. 13. — V. Daubentoni, Leisler, Kuhl, pi. xxv, 2. — V. Nattereri, Kuhl, pi. 23, &c. — Add foreign species, V. emarginatus, Geoff. Ann. Mus. VIII. pi. 46. — V. pictus, L. or the Kirivoula of Java, Seb. I. pi. 56, f. 23.— V. poly thrix, Isid. Geoff. Ann. des Sc. Nat. III. p. 440.— V. levis, Id. ib. &c. t Add V. carolinensis, Geoff. Ann. Mus. VIII. pi. 47. [See Append. I. of Am. Ed.] X Add the Vespertilio of Kuhl, (V. Kuhlii, Natterer), Kuhl, Chauves. d'Allem. p. 55. § Add the Plec. timoriensis, Geoff. — PI. velatus, Isid. Geoff. — PL maugei, Desm. — Plea, cormdus, Fab. — Vesp. megalolis, Rafin. [See Append. II. of Am. Ed.] N. B. — As our plan permits us to class those animals only whose characters we have ascertained either from personal observation or from very coirrplete descriptions and figures, we have been compelled to omit several of the genera of MM. Leach, CARNARIA. 71 Finally, the Nycticees (Rafinesque), have, along with moderate sized ears and the simple muzzle of the Bats, two incisors only in the upper jaw. The known species are from North America*. Galeoihthecus, Pall. The Galeopitheci differ generically from the Bats by the fingers of their hands being furnished with trenchant nails, which are not longer than those of the feet, so that the membrane which occupies the spaces between them, and which is continued as far as the tail, cannot perform any other functions than those of a parachute (a). The canini are denticulated and short like the molars. There are two upper denticulated incisors widely separated from each other ; below there are six, split into narrow strips, like combs, a structure altogether peculiar to this genus. These animals live on trees in the Indian Archipelago, among which they pursue insects, and perhaps birds. If we can judge by the injury the teeth sustain when they become old, they must use fruit also. Their caecum is very large. One species only is well ascertained, the Flying Lemur of Linnaeus. Audeb., Galeeop., pi. 1 and 2. Fur greyish red above, reddish be- low ; spotted . and striped with various shades of grey when young. From the Moluccas and Sunda islands, &c. All the other Carnaria have the mammas situated under the abdomen. FAMILY II. INSECTIVORA.— INSECT EATERS. The animals of this family, like the Cheiroptera, have grinders studded with conical points, and lead most commonly a nocturnal or subterraneous life. Their principal food is Insects, and in cold climates many of them pass the winter in a lethargic (6) state. Unlike the Bats, they have no Rafinesque, & r c. ; and while on this subject, we must observe that there is no family which stands more in need of revision than that of the Bats — a revision from nature and not by compilation. * Vespertilio lasiurus, Schreb., LXIT. B. — V. noveboracensis, Penu. Quadr., pi. 31, fig. 2. — Vcsp. borbonicus, Geoff., Ann. Mus. VIII. pi. 46. £2T {a) Parachute is a French word, which signifies an apparatus to prevent a fall. It strictly applies to a well-known appendage of the air-balloon, which is formed like an umbrella, and is employed by aerial navigators in their descent from the upper regions to the earth, as it is capable, by the resistance which its expansion oilers to the atmosphere, of retarding the fall. — Eng. Ed. ^g" (/)) The lethargy of these hybernating animals has lately received consider- alle scientific attention in this country. Dr. Marshall Hall, after various experiments on Bats {Fespertiliones), the Hedgehog (Erinaceus Europaus), and the Dormouse (Myo.ms Glis), comes to the conclusion that the winter lethargy of these animals is merely a state in which the phenomena characterizing natural sleep only are pre- 12 MAMMALIA. lateral membranes, although they always have clavicles. Their feet are short, and their motions feeble ; the mamma? are placed under the ab- domen, and the penis in a sheath. None of them have a caecum, and in walking they all place the whole sole of the foot on the ground. They differ from each other by the relative position and proportions of their incisors and canines. Some have long incisors in front, followed by other incisors and canines, all even shorter than the molares, a kind of dentition of which the Tar- siers, among the Quadrumana, have already given us an example, and which, to a small extent, approximates these animals to the Rodentia. Others have large separated canines, between which are placed small in- cisors, this being the most usual disposition among the Quadrumana and the Carnaria; and these two dentary arrangements are found in genera, otherwise very much resembling each other in their teguments, the shape of their limbs, and mode of life. Erinaceus, Lin. The body of the Hedgehog is covered with bristles instead of hairs. The skin of their back is furnished with muscles, such as the animal, when bending his head and paws towards the abdomen, can shut himself up within, as in a bag, and present his bristles on all sides to the enemy. The tail is very short, and there are five toes to each foot. There are six incisors in each jaw, the middle ones being the longest, and on each side sented. In ordinary sleep, the Bat, Dormouse, or any other of the lethargic ani- mals, experiences a certain well-marked loss of his respiratory powers; he breathes less, and the temperature of his body diminishes quite perceptibly, so that he can bear with impunity the abstraction of atmospheric air. Now, in the state of hyber- nation, all these symptoms are aggravated — the respiration scarcely goes on at all; and if an animal in this state is placed in a receptacle where the consumption of the gas composing the air can be calculated, it is found that the amount which he ab- sorbs is exceedingly small indeed. The learned gentleman, in these delicate expe- riments, employs a mahogany box, with a glass lid, divided horizontally at its middle part by a fold of strong ribbon. Tbe dimensions of the receptacle should be such as will merely contain the animal. The Bat or Dormouse, being in a state of hyber- nation, should then be placed on the ribbon and inclosed by fixing the lid. A thermometer, which has a cylindrical bulb, should next be passed through an orifice made on purpose in the box, on a level with the ribbon, and should lie beneath and in contact with the upper part of the abdomen of the animal. It should then be left in this situation, and not disturbed; but should be so fixed as that the gas indica- tions should be seen without any interruption of the lethargy; and then those indica- tions, compared with another thermometer hung up in the same apartment, will give the variations of each. In pursuing his investigations, Dr. H. found that the power of animals in a state of hybernation to sustain the want of atmospheric air, enabled a dormant Bat to preserve life for eleven minutes when immersed in water, whereas a Hedgehog, not dormant, died in three minutes when put into the water. A prac- tical distinction of some importance is stated by Dr. Hall between the torpor of ani- mals and their hybernation. The former is an accidental occurrence, arising from a benumbed state of the sentient nerves, and a stiffened condition of the muscles, pro- ceeding directly from cold, and capable of affecting all animals, whilst a defined set of the mammalia only is subject to hybernation, in which state the animal retains its sensibility and power of motion unimpaired. — Eng. Ed. CARNARIA. To three false molars, three molars bristling up ; and a small tuberculated one*. E. europceus, L. ; Buff. VIII. vi. (The Common Hedgehog), has the ears short; common in the woods and hedges; passes the winter in its burrow, whence it issues in the spring with its vesiculaj seminales(a) of an incredible size and complication. To insects, which constitutes its ordinary food, it adds fruit, by which at a cer- tain age its teeth become worn (b). The skin was formerly used to dress hemp. E. auritus, Pall. ; Schreb. CLXIII. (The Long-eared Hedge- hog). Smaller than the preceding; ears as large as the two-thirds of the head, otherwise similar to the europams in form and habits. It is found from the north of the Caspian sea as far as Egypt in- clusively. The Tenrecs, Cuv. (Centenes, Illig). The body of the Tenrec is covered with spines like the Hedgehog. It does not however possess the faculty of rolling itself so completely into a * Pallas has noted as an interesting fact, that the Hedgehog eats hundreds of Cantharides without inconvenience, while a single one produces the most horrible agony in the Dog and the Cat. £3T (a) The vesicular seminales are two membranous sacs, situated beneath the bladder and opening into the urethra or urinary passage. They are receptacles where the secretion, called the semen, is retained. — Eng. Ed. (ST (b) The Hedgehog is very common in the woods, copses, orchards, and thick hedges in England, and its favourite food is beetles, to destroy which it is kept in kitchens. It appears from the account of this creature, given by White, of Selborne, that it can also feed on vegetables: for he states, that he has seen them engaged in the very curious process of devouring the root of a plantain. With the upper man- dible, which is longer than the lower one, they bore under the plant and so eat the root off upwards, leaving the tuft of leaves untouched. But a still more singular fact respecting the food of the Hedgehog, was discovered after an experiment by Professor Buckland, of the University of Oxford. This learned gentleman acciden- tally came to the knowledge of certain circumstances which led him to suspect that Hedgehogs preyed, occasionally at least, on snakes. In order to be satisfied of the truth of his conjecture, he placed the common-ringed snake (coluber natrix), and a hedgehog in a box together. The latter had been bred in an undomesticated state for some time in the Botanical Garden at Oxford, where there was no probability of its having been able to see snakes. At first, the Hedgehog, being rolled up, did not see the snake, when the Professor laid the former on the body of the latter, and in such a way as that the snake was in contact with that part of the ball where the head and tail met. As soon as the snake began to move, the hedgehog started, and open- ing himself up, gave the snake a vigorous bite, and instantly resumed his rolled state. It speedily repeated the bite, and followed it up at the same interval as be- fore with a third bite, by which the back of the snake was broken. The hedgehog then standing by the snake's side, took up and passed through its jaws the whole body of the snake, cracking the bones audibly at every inch. This preparatory pro- cess being completed, the Hedgehog commenced eating the serpent, beginning at the tip of the tail, and, proceeding without interruption, though slowly, consumed it, just as one eats a radish, until about half the victim disappeared. The Hedgehog could not go farther from mere repletion; but it finished the rest of the serpent on the following evening. It is a melancholy fact, that, in many parishes in England, even at the present mo- ment, a bounty is actually paid out of the parish rates for a dead Hedgehog, from the superstitious notion that it sucks the teats of animals. — Eng. En. 74 MAMMALIA. ball: there is no tail; the muzzle is very pointed, and the teeth are very different. There are four or six incisors, and two great canines in each jaw. Behind the canines are one or two small teeth, and four triangular and bristled molars. Three species are found in Madagascar, the first of which has been naturalized in the Isle of France. It is a nocturnal ani- mal, which passes three months of the year in a state of lethargy, although inhabiting the torrid zone. Brugiere even assures us that it is during the greatest beats that they sleep. Erinaceus ecaudatus, L. ; Buff. XII. lvi. (The Tenrec). Co- vered with stiff spines ; only four notched incisors below. It is the largest of the three, and exceeds the common hedgebog in size. Erinaceus setosus, L. ; Buff. XII. lvii. (The Tendrac). The spines more flexible and setaceous ; six notched incisors in each jaw. Erinaceus semi-spinosus. (The radiated Tenrec). Covered with bristles and prickles blended ; striped with yellow and black ; its six incisors and canines are all slender and hooked: size hardly that of a mole*. Cladobates, Fr. Cuv. — Tupaia, Raff. These compose a genus newly established from the Indian Archipelago. Their teeth would greatly resemble those of the hedgehog, were it not that their middle upper incisors are shorter in proportion, that they have four elongated ones in the lower jaw, and that they want the tubercular behind. The animal is covered with hair, has a long shaggy tail, and, contrary to the habits of other insectivora, climbs trees with the agility of a squirrel; the pointed muzzle, however, makes the animal easily distinguishable even at a distance -j\ Sorex, Linn. The shrews are generally small, and covered with hair. Under this, and upon each flank, there is a small band of stiff, thickly set setue, from between which, in the rutting season, oozes an odorous humour, the pro- duct of a peculiar gland \ {a). The two middle upper incisors are hooked * Buff. Suppl. III. pi. 37, has mistaken it for a young Tenrec. Voy. a la Chine, II, p. 146, gives a bad description of the teeth. •f- The banxring; Cladob.javanica, Fr. Cuv.; Tupaia javanica, Horsf. Jav. ; — CI. tana, Fr. Cuv. ; Tup. tana, Horsf. ; — Clad, femigineu, Fr. Cuv.; Tup.ferruginea, Raff. The genus Gymnura of Vigors and Horsfield — Zoolog. Journ. III. pi. 8, appears to ap- proximate to Cladobates by the teeth, and to the Shrew by its pointed snout and scaly tail. There are five unguiculated toes to each foot, and tolerably stiff setae growing among woolly hairs. It can only be properly classed when its anatomy is known. X See Geoff. Mem. du Mus. vol. 1, p. 299. (J3T (a) On each side of the body of this animal there are two sets of glands, one set being destined to secrete milk, whilst the other is intended for very different pur- poses. In the early life of the Shrew, this latter apparatus appears to be merely a longitudinal projection, having no marked characters; but, when the period arrives which exposes the animal to sexual excitement, the projection becomes considerably enlarged, and dotted with innumerable caeca (or minute bladders), which are attached to the body of the gland, and resemble a series of bristles in a brush. The caeca open on the projection of the gland, which has only one duct towards the external surface, and from this the mucus is secreted, whose powerful odour performs so re- markable an agency in the bringing of the male and female together. — Eng. Ed. CARNARIA. 75 and dentated at their base, the lower ones slanting and elongated at their base : five small teeth on each side follow the first, and two only the se- cond. There are moreover in each jaw three bristled molars, and in the upper one, the last is a small tuberculous tooth. The animals live in holes, which they dig in the earth, and seldom leave it till evening; they live on worms and insects, One species only was for a long time known in France ; the Sor. araneus, L. ; Buff. VIII. x, 1. (The Common Shrew, or Shrew Mouse). Grey above; ash-coloured beneath; tail square, and not so long as the body by one-third ; teeth white ; ears naked and exposed; common in fields, meadows, &c. This little animal has been accused of producing a disease in horses by its bite ; the imputation however is false, and arises, perhaps, from the fact, that though cats kill the Shrew, they will not eat it on account of its odour. Daubenton has described the Sor. fodiens, Gm. ; S. Daubentonii, Blumenb. ; Buff. VIII. xi. (The Water Shrew). Rather larger sized than the common one; black above ; white beneath ; tail compressed at the end, and not so long as the body by one-fourth ; the incisors red at the ends ; the ear is surrounded with white, and to a great extent hidden in the hair, and can close itself almost hermetically when the animal dives, while the stiff bristles which fringe its feet give it a facility in swimming, in consequence of which it prefers the banks of rivulets. Several Shrews have been observed in Europe, which differ in some respects from the preceding ones ; but as in this genus the age and season materially affect the colours of the fur, it is by no means cer- tain that they are all constant species* Other countries also have their own, the most remarkable of which is the S. myosorus, Pall., Act. Petrop. 1781, part II, pi. 4; Mus musquee de Vlnde, Buff. Supp. VII. 71. (The Rat-tailed Shrew). In its form and colour it resembles our common Shrew, and also has its large naked ears ; but the tail is round, furnished only with hairs, plainly scattered, and is almost as large as that of our long-tailed field-mouse. It gives out a strong musky scent, which impregnates every thing it touches. It is found throughout India, and part of Africa, and is one of the animals the ancient Egyptians were in the habit of embalming -j~. * The S. leucodon, Sclireb. 159, D, does not appear to me to differ from the com- mon Shrew. I strongly suspect the S. tetragonurus and constrictus, Herm., Sclireb. 159, B and C, or Geoff. Ann. Mus. XVII. pi. 3, f. 3, and pi. 3, f. 1, and even the S. remifer, Geoff. Ann. Mus. XVII. pi. 2, f. 1, to be aged Water-Shrews; the remifer particularly, whose belly is sometimes white, sometimes black; the S. lineatus, Geoff, ib. 181, is an accidental variety of the tetragonurus arising from age. The Sorex minutus, Laxmann, Schreb. 161, B, is merely a mutilated specimen of the S. pig- ments, Pall. Such is not the case however with the S. etruscus, Savi, which is but half the size of our common species, is blackish, has naked ears, white muzzle and paws, round tail, &c. It is a true and distinct species. f I consider the S. myomrus Pall, and Geoff. Ann. du Mus. XVII. pi. 3, f. 2; the S. capensis, Id. ib. pi. ii. f. 2; the S. indicus, Id. Mem. du Mus. I, pi. 15, f. 1, as old spc- 76 mammalia. Mygale, Cuv. The Desmans differ from the shrews by two very small teeth placed be- tween the two great incisors of the lower jaw, and in their two upper in- cisors, which are triangular and flattened. Behind these incisors are six or seven small teeth and four bristled molars. Their muscle is drawn out into a little flexible proboscis, which they keep constantly in motion. Their long tail, scaly, and flattened on the sides, and their five-fingered feet all united by membranes, evidently proclaim them to be aquatic ani- mals. Their eyes are very small, and they have no external ears. Sorex moschatus, L. ; Buff. X. 1.; Pall. Act. Petrop. 1781, part II, pi. 5. (The Russian Musk Rat). Nearly as large as a shrew; above, blackish; beneath, whitish; tail not so long as the body by one-fourth. Very common along the rivers and lakes of southern Russia, where it lives on worms, the larva? of insects, and particularly on leeches, which, by means of its flexible snout, it easily withdraws from the mud. Its burrow, which is made in the beach, commences under water, and ascends to such a height as to be above its level in the greatest floods. This animal never comes voluntarily on shore, but numbers of them are taken in the nets of the fishermen. Its musky odour arises from a kind of pomatum that is secreted in small follicles under the tail, and it is so powerful as to be communicated to the flesh of the pike, which feeds on the musk rat. A small species of this genus is found in the rivulets of the Pyre- nees, whose tail is longer than its body, which M. Geoff, has made known, Ann. du Mus. torn. XVII. pi. iv. f. 1, Myg. pyrendica, H. Chrysochloris, Lacep. Have, like the preceding genus, two incisors above, and four below; but their grinders are long, distinct, and almost all shaped like triangular prisms. Their muzzle is short, broad, and recurved, and their fore-feet have only three nails, of which the external, being very large, extremely arcuated and pointed, serves them as a powerful instrument for excavating and piercing the earth ; the others regularly decrease in size. The hind feet have five of an ordinary size. They are subterraneous animals, whose mode of life is similar to that of moles. To enable them to dig the better, their fore-arm is supported by a third bone placed under the cubitus. C. asiatieus; Talpa asiatica, L. ; Schreb. CLVII; and better, Brown, 111. XLV. (The Golden Mole). A little smaller than the European mole ; no apparent tail ; is the only quadruped known that presents any appearance of those splendid metallic tints which bright- cimens or varieties of one and the same species, to which I also refer the S. giganteua, Isid. Geoff. Mem. du Mus. XV. pi. 4, fig. 3; perhaps even the S.flavescens, Isid. Geoff", ib. Seba figures it, Mus. I. pi. 31, f. 7 and 11— pi. 63, fig. 5, and the white variety, I. pi. 47, f. 4. — Add the S. murinus, Lin. of Java, of the size of a mouse; grey; ears naked; tail round and nearly as long as the body.— The S. brevicaudus, Say, from North America; blackish, ears concealed, tail one-fourth the length of the body. — S. parvus. Id. with naked ears. — The S. suaveolens, Pall., and the other species pointed out by him in his Zoography of Kussia. This genus needs revision as much as that of the Bats. CARNARIA. 77 en and adorn so many birds, fishes, and insects. Its fur is a green, changing to a copper or bronze colour; there is no conch to the ear, and no eyes can be discovered*. Talpa, Lin. The moles are universally known by their subterranean life, and by their form, which is eminently fitted for their mode of existence. A very short arm attached by a long scapula, supported by a powerful clavicle, furnish- ed with enormous muscles, sustains an extremely large hand, the palm of which is always directed either outwards or backwards ; the lower edge of this hand is trenchant, the fingers are scarcely perceptible, but the nails in which they terminate are long, flat, strong, and sharp. Such is the in- strument employed by the mole to tear the earth, and throw it backward. Its sternum, like that of birds and bats, has a process which gives to the pectoral muscles the large size that is required for their functions. To pierce and raise up the earth, it makes use of its long pointed head, whose muzzle is armed at its extremity with a peculiar little bone, and its cervical muscles are extremely powerful. There is even a special bone in the cer- vical ligament. It has but little power behind, and moves as slowly above ground as it advances rapidly under it. Its sense of hearing is very acute, and the tympanum very large, although there is no external ear ; its eyes are so small, and so hidden by the hair, that for a long time their exist- ence was positively denied. In the genital organs there is this peculiarity — the bones of the pubis are not united, a circumstance which permits it to produce tolerably large young ones, notwithstanding the narrowness of the pelvis. The urethra of the female passes through the clitoris. She has six teats. The jaws are weak, and the food consists of worms, insects, and some soft roots. There are six incisors above, and eight below. The canines have two roots, which causes them to partake of the nature of false molars; behind, there are four false molars above and three below, after which are three bristled molars. T. europcea, L. ; Buff. VIII. xii. (The Common Mole). Point- ed muzzle ; fur thin and black ; individuals are found white, fawn- coloured, and piebald. This is an animal which is found very incon- venient by the havoc which he makes in gardens and other cultivated places. This species, according to Dr. Harlan, is also found in North America (a). * The Red Mole of America, Sebay, pi. xxxii. f. 1, {Talpa rubra, Lin.) is most pro- bably a Chryso asiaticus, drawn from a dried specimen of that species, for then the fur appears purple; the tucan of Fernandez, App. XXIV, which is considered as syno- nymous with it. (J3T (a) We have the authority of Dr. H. M'Murtrie against the existence of the common Mole in America. That which obtains the name of the common Mole in the United States is the Scalops Aqualicus, described in the succeeding page. — Eng. Ed. g^f The common Mole, at the period when it builds its nest, (about the beginning of February), and brings forth its young, is an object peculiarly deserving the attention of the naturalist. The nest is always in a cavity formed in the midst of the hillocks, which are so frequently to be met with in the fields, and well known under the title 78 MAMMALIA. M. Savi has found a Mole in the Apennines that is perfectly blind, although otherwise similar to the common one; he calls it Talpa cceca. CONDYLURA, Ill'lg. The Condylures seem to combine the two kinds of dentition of the insec- tivora. In the upper jaw are two large triangular incisors, two extremely small and slender ones, and on each side a strong canine. In the lower one are four incisors slanting forwards, and a pointed but small canine. The superior false molars are triangular, and separated, the inferior trench- ant and denticulated. In their feet and the whole of their exterior they resemble the Mole, but their tail is longer, and what more particularly distinguishes them from the former is, that their nostrils are surrounded with little moveable carti- laginous points, which, when they separate, radiate like a kind of star. One species particularly is found in North America — Sorex cris- tatus, L*. (The Radiated Mole). Similar to the Mole of Europe, the nose excepted, but having a tail more than double the length of that of the latter. SCALOPS, CiW. The Scalops have the teeth very similar to those of the Desmans, except that the small or false molars are less numerous ; the muzzle is simply pointed, like that of the Shrews; their hands are widened, armed with strong nails fitted to excavate the earth, and exactly similar to those of Moles : in fact, their mode of life is the same ; their eyes are equally as small, and their ears quite as much hidden. The only species known is the *S". aquations; Sorex aquations, L. ; Schreb. CLVIII. (The Ca- nadian Scalop). It appears to inhabit a great part of North America, along rivers, &c. Its external resemblance to the common Mole of Europe is so great, that it is easy to mistake the one for the other. * This is the Condylura of Illiger, but the characters he indicates, taken from the figure of La Faille, copied Buif. Supp. VI. xxxvi, 1, and on which he composed the name of the genus, are false. M. Desmarets was the first who correctly described the teetli of this animal. Dr. Harlan describes a species, Cor.d. macroura, which has but very short points about the nostrils, and a scaly compressed tail. He associates with it, as a third spe- cies, the Talp. longicaudata, Penn. Hist. No. 443, which he appears however not to have seen. of mole hills. Moss forms the principal lining of the nest. One of the most curious phenomena presented by the Mole at this period is the process of skinning a worm. The integument of the victim is stripped from end to end, and then the contents are squeezed out by pressure on the part of the mole. The subterranean burrows formed by these animals are mostly connected by avenues with the chamber where the nest is made; and in going to or from this chamber, the mole passes through a series of these avenues, where, it is supposed, that several means, known only to the contriver, are employed for catching worms, beetles, grubs, &c. The inconvenience to which Cuvier alludes, as being produced chiefly in cultivated places, consists, for the most part, in the loosening of the earth round the roots of plants, which always attends their operations in searching for food. — Enc. Ed. CARXARIA. 7<) FAMILY III. CARNIVORA. Although the term carnivorous (Carnassiers) is applicable to all un- guiculated animals, not quadrumanate, that have three sorts of teeth, inas- much as they all feed more or less on animal aliment, there are, however, many of them, the two preceding families especially, which are compelled by weakness, and the conical tubercles of their grinders, to live almost en- tirely on insects. It is in the present family that the sanguinary appetite for flesh is joined to the force necessary to obtain it. There are always four stout, long, and separated canines, between which are six incisors in each jaw, the root of the second of the lower ones being placed a little more inwards than the others. The molars are either wholly trenchant, or blended merely with blunted tuberculous parts, but they are not bristled with conical points. These animals are so much the more exclusively carnivorous, as their teeth are the more completely trenchant, and the proportions of their re- gimen may be calculated from the extent of the tuberculous surface of their teeth, as compared with that portion which is trenchant. The Bears, which can live altogether on vegetables, have nearly all their teeth tuber- culated. The anterior molars are the most trenchant; next comes a molar larger than the others, usually furnished with a larger or smaller tuberculous heel; and behind it one or two small teeth, that are perfectly fiat. It is also with these small teeth in the back part of the mouth that the dog chews the grass he sometimes swallows. We shall call, with M. Fr. Cu- vier, this large upper molar, and its corresponding one below, lacerators (carnassiers); the anterior pointed ones, false molars; and the posterior blunted ones, tuberculous teeth. It is easy to conceive that those genera which have the fewest false mo- lars, and whose jaws are the shortest, are those best adapted for biting. It is upon these differences that the genera can be most safely esta- blished. It is necessary, however, that the consideration of the hind foot should be added to them. Several genera, like those of the two preceding families, in walking, or when they stand erect, place the whole sole of the foot on the ground, a fact proved by the total want of hair on every part of the sole. Others, and by far the greater number, walk only on the ends of the toes, by raising up at the same time the tarsus. They are much swifter; 80 MAMMALIA. and to this leading distinction are added many others in the habits, and even in the internal conformation. In both the substitute for the clavicle is a mere bony rudiment suspended in the flesh. The Plantigrada Form this first tribe, which walks on the whole sole of the foot, a cir- cumstance which gives them a greater facility in balancing themselves upon their hinder feet. They partake of the slowness and nocturnal life of the Insectivora, and, like them, have no caecum : most of those that in- habit cold countries pass the winter in a state of lethargy. They all have five toes to each foot. Ursus, Linn. The Bears have three large molars on each side* in each jaw, altogether tuberculous, and of which the posterior upper, and anterior upper, are the longest. They are preceded by a tooth a little more trenchant, which is one of the lacerators of this genus, and by a variable number of very small false ' .lolars, which are sometimes shed at a very early period. This almost frugivorous sort of dentition is the reason why, notwithstanding their great strength, they seldom eat flesh, unless from necessity. They are large stout-bodied animals, with thick limbs, and a very short tail : the cartilage of the nose is elongated and moveable. They excavate dens or construct huts, in which they pass the winter in a state of somno- lency more or less profound, and without food. It is in these retreats that the female brings forth. The species are not easily distinguished by apparent characters. We have the U. arctos, L., Buff. VIII. xxxi. (The Brown or common Bear of Europe). Forehead convex : fur brown, more or less woolly when young, and growing smoother with age. Some of them are greyish, others almost yellow, and a third kind is brown, with shades border- ing on silver. The relative height of their legs is equally variable, and all without any fixed relation to age or sex. They have most com- monly, when young, a whitish collar, which, in some varieties, re- mains for a longer or shorter period, and even for life. This animal inhabits the lofty mountains, and great forests throughout Europe, and of a great part of Asia ; the coupling season is in June, and the young are produced in January. It sometimes lodges very high up in trees ; when young its flesh is esteemed a delicacy — the paws are considered good at all ages. It is thought that the Blade Bear of Europe is a distinct species : those which have been described as such had a flat forehead, and the fur woolly and blackish ; their origin, however, does not appear to us to be very authentic : \. * We shall hereafter omit the repetition of the words " on each side," &c, it be- ing understood that we speak of the molars on one side only, those of the other being the same. \ We are not yet satisfied that the Grisly Bear of North America differs specifi- cally from the Brown Bear of Europe. P. S. Since the above note was written, General La Fayette has presented a Grisly Bear to the Menagerie du Jardin du Roi. In form and hair, some shades of colour- CARNARIA. 81 U. americanus, Gm.; Fr. Cuv. Mammil'. ; Schreb. pi. 141, B. (The North American Black Bear). A very distinct species, with a flat forehead, smooth and black fur, and fawn-coloured muzzle. We have always found the small teeth behind the canines more numerous in this bear, than in the European species. Individuals have been seen that were entirely fawn-coloured. Its usual food is wild fruits ; it devastates the fields, and, where fish is abundant, proceeds to the shores for the purpose of catching it. It is only for want of other aliment that it attacks quadrupeds. The flesh is held in great esteem. There is another Black Bear found in the Cordilleras, with a white throat and muzzle, and large fawn-coloured eyebrows, that unite on the forehead— U. ornatus, Fr. Cuv. Mammif. The East Indies also produce several bears of a black colour, such as the U. malaianus, Horsf. Java. (The Malay Bear). From the Pe- ninsula beyond the Ganges and the islands of the Straits of Sunda. Smooth; black; fawn-coloured muzzle; a heart-shaped spot of the same colour on the breast. It is very injurious to the cocoa nut trees, which it climbs in order to devour their tops and drink the milk of the fruit. U. thibetanus, Fr. Cuv. Mammif. (The Thibet Bear). Black ; the under lip and a large mark in the form of a Y in white on the breast; profile straighter and claws weaker. From the mountains in the north of India. But the most remarkable of these Bears of India is the U. labiatus, Blain. ; IS Ours jongleur, Fred. Cuv. Mammif.; U. longirostris, Tied. (The Thick-lipped Bear). The cartilage of the nose dilated; the tip of the under lip elongated, both being moveable ; when old, very thick bushy hairs round the head. The facility with which the incisors are lost, occasioned it for a long time to be considered as a Sloth*. It is black; the muzzle and tips of the paws fawn-coloured or whitish, and a half collar or spot in the form of a Y under the neck and breast. This animal is a favourite with the Indian jugglers, which they lead about on account of its deformity. U. maritimus, L.; Cuv. Menag. du Mus., 8vo., p. 68; copied, Schreb. pi. clxi. (The Polar Bear). This is another species, very distinguishable by its long and flattened head and its white and ing excepted, it closely resembles the Brown Bear; its nails, however, are much longer and more trenchant. It appears to be a distinct species. M. Horsfield, Lin. Trans. XV. 332, describes a bear from Nepaul, of a light bay colour, whose nails are less trenchant than those of the other bears of India, and which appears to him a distinct species. I have neglected stating in the text that we have recovered many fossil bones of lost species of bears, the most remarkable of which are the U. spelaus, Blumenb., with a rounded forehead, and of a very large size; and the U. cultridens, Cuv. See the fourth vol. of my " Ossemens Fossiles." * It is the Brady-pus ursinus of Shaw, and the genus Prochilus, Illig. See Jour, de Phys. of 1792, vol. xl. p. 136. VOL. I. G 82 MAMMALIA. smooth fur. It pursues seals and other marine animals. Exagger- ated accounts of its ferocity have rendered it highly celebrated (a). ({^* (a) In the Zoological Gardens, Regent's Park, may be seen the following Bears: — Sloth Bears, Ursics labiatus, Blain., male and female. They are the species usually exhibited by the Indian jugglers, and have been found milder in their dispositions than most of the other Bears. They are from India. Malay Bear, or Malayan Sun Bear, Ursus Malay anus, Horsfield. This bear was brought from Sumatra, and is found only in places near the equator. It feeds chiefly on vegetables, and is particularly fond of the young shoots of the cocoa nut tree. It is likewise fond of honey, and the tongue, as may be seen, is elongated, so as to adapt it admirably to the process of lapping. Spectacled Bear, Ursus ornatus of F. Cuvier. This Bear is a great curiosity; it is a native of the mountains of South America, and has been but recently described; this is the first of the species that has been seen in this country. American Black Bear, Ursus Americamis. This Bear is from North America, and feeds on vegetable roots and fruits; and, in its native state, resorts to the sea coasts for the purpose of consuming fish. It appears to be smaller in size than the Euro- pean bear. Cinnamon Bears, called also Chocolate Bears, are considered only as varieties of the Black Bears. The former, however, are distinguished from the latter by the marked difference of their habitats ; for, whilst the Black Bears keep to close and woody districts, the Cinnamon Bears live in the open and upland grounds; the latter are described also as more powerful and vicious than the others in the native state. The European Brown Bear, Ursus arctos, Linnaeus. This bear is from Siberia, and its diet is altogether vegetable. The flesh of the young bears of this species is eatable. Siberian Bear, Ursus collaris, F. Cuvier. This bear is particularly distinguished by the white collar around his neck. But some naturalists doubt if the animal be not a mere variety of the Brown Bear. Grisly Bear, Ursus ferox, Lewis and Clarke. This is an object of great curiosity, inasmuch, as in all probability it is the largest and most powerful of the bear tribe; it is certainly the most ferocious, and is described as possessing extraordinary tenacity of life. This specimen was brought from the rocky mountains in North America, about the year 1813, to England, and was placed in the menagerie of the Tower. It formed part of the munificent present, made by his present Majesty on his accession, to the Zoological Society. White or Polar Bear, Ursus maritimus, Gmelin. This is a remarkable female spe- cimen of the White Bear, which is found in no other part of the world except the coldest of the northern regions. This Bear appeared formerly to be the largest; but its size was greatly misrepresented by the older navigators. Captain Parry, in his north-west expedition, has not met with one which exceeded seven or eight feet. This bear preserves uniformly a white colour on every part of its external surface, except on the naked end of the snout, the lips, and the margins of the eyelids and claws. There is no doubt whatever, that the female Polar Bears, when pregnant, are subject to undergo hybernation ; and, it is j>robable, that the males merely suffer from the effects of torpidity, the nature of which will be found explained in a previousnote to the Hedge- hog. The distinction pointed out in that note between hybernation and torpidity affords a medium, whereby the contrary opinions of naturalists on this point may be recon- ciled. This Bear is chiefly hunted for its hide and fur. The accounts which we pos- sess of the Brown Bear, so common in the Scandinavian forests, are the result of a persevering scientific study of the habits of these animals. They consist, according to some, of two varieties of the Ursus arctos, the large Bear, or Bear of prey, in Swedish, Slag-Bjorn, which lives indiscriminately on animal and vegetable substances, is one; and a smaller species, in Swedish, Myr Bjorn, which subsists en- tirely on ants or vegetable substances, forms the second. Other naturalists, however, are of opinion that there is only one real species in Scandinavia, and that it is omni- vorous, feeding on cattle indiscriminately, and on roots, leaves, small branches and berries of all sorts. One remarkable habit of this be<.^t is, that about October every year he ceases to feed for the winter season; his stomach and intestines then become CARNAR1A. 88 Procyon, Storr. The Ratons or Raccoons have three back tuberculous molars, the su- perior of which are nearly square, and three pointed false molars in front, forming a continuous series to the canines, which are straight and com- pressed. Their tail is long, but the remainder of the exterior is that of a bear in miniature. They rest the whole sole of the foot on the ground only when they stand still; when they walk they raise the heel. P. lotor; Ursus lotor, L. ; Mapach of the Mexicans ; Buff. VIII. xliii. (The Raccoon). Greyish brown ; muzzle white ; a brown streak across the eyes ; tail marked with brown and white rings. This animal is about the size of a Badger, is easily tamed, and re- markable for a singular habit of eating nothing without having pre- viously dipped it in water. From North America — lives on eggs, birds, &c. P. cancrivorus; Ursus cancriv. L. ; Buff. Supp. VI. xxxii. (The Raccoon Crab-eater). A uniform light ash-brown ; the rings on the tail less distinct. From South America. Ailuhus, Fred. Cuv. The Panda appears to approximate to the Raccoon by its canines, and what is known of its other teeth ; with this exception, that it has only one false molar. The head is short ; tail long ; walk plantigrade ; five toes, with half retractile nails*. One species only is known, the A. refulgens, Fred. Cuv. Mammif. ; Hardwick, Linn. Trans. XV. p. 161. (The Shining Panda). Size of a large cat ; fur soft and thickly set ; above of the most brilliant cinnamon red ; behind more fawn-coloured; beneath of a deep black. The head is whitish, and the tail marked with brown rings. This most beautiful of all known quadrupeds, and which inhabits the mountains of the north of India, was sent to Europe by my son-in-law the late M. Alfred du Vaucel. Ictides, Valenciennes. The Benturong is somewhat related to the Raccoon by its teeth ; but the three upper back molars are much smaller and less tuberculous ; and this is especially the case in the last one in each jaw, which is very small * General Hardwick has described the upper teeth of the Panda, Lin. Trans. XV. pi. ii. There are four square and tuberculous grinders, and one false trenchant mo- lar in front, at a short distance from the canine. quite empty, and collapse into a small compass within the abdomen, whilst the ex • tremity of the last bowel is blocked up by a piece of hard wadding, called in Sweden Tappen. In November, he retires to his den, which is usually prepared before-hand, and here he undergoes the state of hybernation. About the middle of the following April, the bear quits his den, voids the tappen, and bounds with fresh vigour into the forest in search of food. The tappen, when chemically examined, has been found to consist of the following ingredients : brown resin, green volatile oil, colouring matter of leaves, fat, starch, woody matter, with the acids and salts of trees in the Scandina- vian forests. We are unable to find any account of the substance called Tap/ 1 n in the most complete systematic works on the subjects of comparative anatomy and ani- mal secretions. We have carefully consulted the great work of Berzelius, himself a Swede; but we can find no allusion to the substance amidst the vast details which he gives of the secretions of the Mammalia. — Eng. En. G 2 St mammal;.'.. and nearly simple. It is covered with long hair, and has a tuft of it at each ear. The tail is long, hairy, and has a propensity to curl, as if pre- hensile. This animal is also one of those from India, for the knowledge of which we are indebted to the late M. du Vaucel. One species is the let. albifrons, Fr. Cuv., Ann. des Sc. Nat. IV. pi. 1. Grey; tail and sides of the muzzle black; size that of a large cat. From Bootan. let. ater, Fr. Cuv. Mammif. Black; muzzle whitish; size that of a stout dog. From Malacca*. Nasua, Storr. The Coatis, to the teeth, tail, nocturnal habit, and slow dragging gait of the Raccoon, add a singularly elongated and flexible snout. The feet are semi-palmate, notwithstanding which they climb trees. Their long nails are used for digging. They inhabit the warm climates of America, and their diet is nearly the same as that of the Marten of Europe. Viverra nasua, L. ; Buff. VIII, xlviii. (The Red Coati). Red- dish fawn colour ; muzzle brown; tail with brown rings. Viv. narica, L. ; Buff. VIII, xlviii. (The Brown Coati). Brown, white spots over the eye and snout. This is, perhaps, the only proper place for the singular genus of the Kinkajous or Potto, Cuv. — Cercoleptes, Illig. — which, to a planti- grade walk, adds a long prehensile tail, like that of the Sapajous, a short muzzle, a slender and extensible tongue, two pointed grinders before, and three tuberculous ones behind. Only one species is known, Viverra caudivolvula, Gm. ; Buff. Supp. III. 4; and better, Fr. Cuv. Mammif. From the warm parts of America, and from some of the great Antilles, where it is called Potto; size of a Polecat; hair woolly, and of a grey or yellowish brown; habits nocturnal, of a mild disposition, and lives on fruit, milk, honey, blood, &c. Meles, Storr. The Badgers, which Linnaeus placed with the Raccoons, among the Bears, have a very small tooth behind the canine, then two pointed molars, fol- lowed in the upper jaw by one that we begin to recognise as a lacerator, from the trenchant vestige it exhibits on its outer side; behind this is a square tuberculous one, the largest of all. Below, the penultimate begins to shew a resemblance to the inferior carnivorous teeth ; but, as there are two tubercles on its internal border, as elevated as its trenchant edge, it acts as a tuberculous one ; the last below is very small. These are animals with a rampant walk and nocturnal habits, like the preceding, whose tail is short, fingers considerably enveloped in the skin, and which are further conspicuously distinguished by a pouch situated be- neath the tail, and from which a greasy and fetid secretion oozes out. Their very elongated fore nails render them exceedingly dexterous in dig- ging the ground. M. europcea; Ursus meles, L. ; Buff. VII, vii. (The European Badger). Greyish above, black beneath, a blackish band on each * Add Vlrlide dore, Fred. Cuv. CARNARIA. 85 side of the head. The American Badger {Mel. hudsonius) is not very different. Gulo, Storr. Linnaeus also placed the Gluttons among the bears ; but they approximate much nearer to the weasels in their teeth as well as in their habits ; the only relation they have to the former consisting in their plantigrade movement. They have three false molars above, and four below; in front, the carnivorous one, which is well characterized; and behind it, a small tuberculous one — the upper being more broad than long. Their su- perior carnivorous tooth has only one small tubercle on the inner side, and, in fact, the whole dental system is nearly the same as that of the weasels. The tail is of a middling size, with a plait or fold beneath, in place of a sac, and the port of the animal is very similar to that of the badger. The most celebrated species is the Glutton of the north, the Ursus gulo, L. ; Buff. Supp. Ill, xlviii. (The Common Glutton or Rosso- mak of Russia). About the size of the Badger; usually of a fine deep maronne colour, with a disk on the back, of a darker brown ; sometimes, however, the shades are lighter. It inhabits the most glacial regions of the North, is considered very sanguinary and fe- rocious, hunts during the night, does not become torpid during the winter, and masters the largest animals by leaping upon them from trees. Its voracity has been ridiculously exaggerated by some au- thors. The Wolverene of North America {Ursus luscus, Lin. Edw. 103) does not appear to differ from it in any constant character; its colours, however, are generally lighter. Hot climates produce some species which can only be placed near the Gluttons, as they differ from them merely in having one false molar less in each jaw, and in a long tail. Such are the animals termed by the in- habitants of South America Ferrets, which, having the teeth of our Fer- rets and Polecats, have, in fact, similar habits ; they are distinguished from them, however, by their plantigrade movement. Viverra vittata, L. ; Buff. Supp. VIII, xxiii and xxv. (The Gri- son). Black ; top of the head and neck grey ; a white band, reach- v ing from the forehead to the shoulders. Mustela barbara, L. ; Buff. Supp. VII, lx. (The Taira). Brown ; top of the head grey ; a large white spot under the throat. These two animals are found in all the warm climates of America, and smell strongly of musk. Their feet are somewhat palmated, and it appears they have sometimes been taken for otters*. Ratelus, F. C. The Ratels have a false molar in each jaw less than the Grison, and their upper tuberculous tooth is but slightly developed, so that in the teeth * It is supposed, by the description given by Marcgrave of his cariqueibeiu, which name Button has applied to his saricovienne, Vol. XIII. p. 319, that he meant to speak of the Taira. 8(5 MAMMALIA. they approach the Cat, while their whole exterior is that of the Grison or Badger. The legs are short; feet plantigrade, and five toes to each; nails very strong, &c. &c. One species only is known, the Viverra mellivora, Sparm. ; and Viv. eapensis, Schreh. pi. 125. (The Ratel or Honey Weasel). Size of the European Badger; grey ahove; hlack beneath, with a white line between these two colours ; sometimes it is nearly all white above. It inhabits the Cape of Good Hope, and digs up the earth with its long fore-claws, in search of the honey-combs of the wild bees. The DlGITIGRADA form the second tribe of the Carnivora — that which walks on the ends of the toes. In the first subdivision there is only one tuberculous tooth behind the upper carnivorous one ; these animals, on account of the length of their body, and the shortness of their feet, which permit them to pass through the smallest openings, have been styled vermiform. Like the preceding ones, they have no caecum, but do not fall into a lethargy during the winter. Linnaeus placed them all in one genus, that of Mustela, Lin. Or the Weasels, which we will divide into four sub-genera. Putorius, Cuv. The Polecats are the most sanguinary of all; the lower carnivorous tooth has no inner tubercle, and the superior tuberculous one is more broad than long; there are only two false molars above and three below. These animals are externally recognised by their muzzle, which is shorter and thicker than that of the Weasel. They all diffuse an infectious odour. Mustela putorius, L. ; Buff. VII. xxiii. (The Common Polecat). Brown; flanks yellowish; white spots on the head; the terror of poultry-yards and warrens. M. furo, L. ; Buff. VII. xxv. and xxvi. (The Ferret). Yellow- ish, with rose-coloured eyes, and is perhaps a mere variety of the Polecat. It is only found in France in a domesticated state, and is employed to ferret out the rabbits from their holes. It comes from Spain and Barbary. M. sarmatica; The Perouasca; Pall. Spic. Zool. XIV. iv. 1; Schreb. CXXXII. (The Sarmatian Weasel). Brown; everywhere spotted with yellow and wbite. So beautifully is the skin mottled, that it is in high request among furriers. It is found throughout all southern Russia, Asia Minor, and the coast of the northern rivers of the Caspian sea. M. sibirica, Pall. Spic. Zool. XIV. iv. 2. (The Siberian Pole- cat). A uniform light fawn colour ; nose and circumference of the eyes brown ; end of the muzzle and the under part of the lower jaw white. CARNARIA. 87 It is also to this subdivision that we must refer two small Euro- pean species — M. vulgaris, L. ; Buff. VII, xxix, 1. (The Weasel). Of a uni- form red; and the M. erminea, L.; Buff. VII, xxix, 2, and xxxi, 1. (The Stoat or Ermine). Red in summer, white in winter ; end of the tail always black. The winter skin is one of the best known furs. We should also place near it the M. lutreola, Pall. Spic. Zool. XI. 1; Leche, Stock. Mem. 1739, pi. xi; Schreb. CXXVII. (The Mink or Norek, or Polecat). It frequents the shores of rivers, &c, in the north and east of Europe from the Arctic Ocean to the Black Sea, and lives on frogs and crabs. The feet are slightly palmated at the base of the toes, but the teeth and round tail approximate it nearer to the Polecat than the Otter. It is of a reddish brown ; the circumference of the lips and the under part of the jaw white ; it exhales a musky odour, and its fur is very beautiful. The above animal is considered by some to be the same as the Polecat of the North American rivers, to which the name of Mink has been transferred, whose feet are likewise semi-palmated ; but the only white about it is on the point of the chin, and sometimes a nar- row line under the throat — it is a different species*. Warm climates also have their Polecats or Weasels. Put. nudipes, Fred. Cuv. Mammif. (The Javanese Polecat). Golden-yellow; head and tip of the tail white. Put. africanus, Desm. (The African Polecat). Reddish fawn colour above ; yellowish white below ; a red band reaching longitu- dinally along the middle of the belly from the fore to the hind legs. Put. striatus, Cuv. (The Striated Madagascar Ferret). Size of the European Weasel; reddish brown, with five longitudinal white stripes; the under part and the tail nearly all white. Put. zorilla; Zorille, Buff. ; Viverra zorilla, Gm. ; Buff. XIII. xl. 1 . (The Zorilla, or Cape Polecat). Irregularly striped with black and white ; an animal that has been so far confounded with the mephitic weasels as to receive the name of Zorillo, or little Fox, which the Spaniards have applied to those fetid American animals. It approaches them in its claws, which are fitted for digging, but in every thing else resembles the Polecats. They indicate a subter- raneous habit, which might induce us to separate it from the other species. Mustela, Cuv. The true Weasels differ from the Polecats in having an additional false molar above and below, and in the existence of a small internal tubercle * When this page was written, I had no other knowledge of the Norek, or Mink of Europe, than what the description of Pallas afforded me. Having since then procured some specimens, I have ascertained that the white about the jaws is not permanent, and that very frequently the only white to be seen is at the end of the lower jaw, as in the American Mink. I now think they are both one species. 88 MAMMALIA. on their inferior carnivorous tooth, two characters which somewhat diminish the cruelty of their nature. There are two species in Europe closely allied to each other, the M. martes, L. ; Buff. VII. xviii. (The Pine Marten). Brown; a yellow spot under the throat. Inhabits the woods. M. foina, L. ; Buff. VII. xviii. (The Common Marten). Brown ; the whole under part of the throat and neck white. Inhabits houses. Both species are very destructive. Siberia produces the M. zibellina, Pall. Spic. Zool. XIV. iii. 2 ; Schreb. CXXXVI. (The Sable). So celebrated for its rich fur; brown, spotted with grey about the head, and distinguished from the preceding ones by the extension of the hair to the under surface of the toes. It inha- bits the coldest mountains, and the hunting to obtain it, in the midst of winter and tremendous snows, is the most painful with which we arc acquainted. It is to the pursuit of this animal that we owe the discovery of the eastern countries of Siberia(a). North America also possesses several Martens indicated by na- turalists and travellers, under the indefinite names of Pekan, Viso?i, Mink, &c. One of them, the White Vison of the furriers, Mus. leutreoce- phala, Harl., has as hairy feet and almost as soft a fur as the Sable, but is of a light fawn colour, and almost white about the head. That which we call the Pekan; Must, canadensis, Gm., and which comes from Canada and the United States, is of a brownish colour, mixed with white on the head, neck, shoulders and top of the back; nose, crupper, tail and limbs blackish*. Mephitis, Cuv. The Skunk, like the Polecat, has two false molars above and three be- low, but the superior tuberculous one is very large, and as long as it is broad, and the inferior carnivorous has two tubercles on its internal side ; circumstances which ally it to the Badger just as the Polecat approximates to the Grison and Glutton. Independently of this, the anterior nails of the Skunk, like those of the Badger, are long and fitted for digging; they are moreover semi-plantigrade, and the resemblance extends even to the distribution of their colours. Amongst this family so remarkable for its stench, the Skunks are distinguished by a sort of stench far exceeding that of the remaining species. * It is the Pekan of Daubenton, but it has not always the white under the throat. There are several other species of Polecats or of Martens indicated by MM. Molina, Humboldt, and Harlan; but they require re-examination (6). 1& (a) The "painful" task here alluded to was imposed exclusively, during the more barbarous periods of Russian tyranny, on the unhappy exiles who were trans- ported on the most unjust pretences to the wilds of Siberia. These persons were under an obligation to furnish, within a given time, a defined number of sable furs. — Eng. Ed. @5F (b) A specimen of the Pekan or Fisher Marten may be seen in the Zoological Gardens, Regent's Park. Its name of Fisher is not appropriate, as it does not eat fish, but pursues the same prey as the Pine Marten. — Eng. Ed. CARNARIA. 89 Skunks are generally striated with white stripes on a black ground, but the number of stripes appears to vary in the same species. The most common species of North America is the M. putorius; Viverra putor, Gm. ; Catesb. Carol. II. lxii. Schreb. CXXII. (The American Skunk — the Fitchet of Pennant). Black, with stripes of white, larger or smaller, and more or less nu- merous ; the tail is black, and the tip white. The odour it produces resembles that of the polecat, mingled with a strong smell of garlic — nothing is more nauseous. It would seem that in South America the species most usually encountered has a white tail. The stripes on the back sometimes occupy its whole breadth ; it is the Viverra mephitis, Gm.; Buff. XIII. xxxix, or the Chinche*. We may make a distinct subgenus of the Mydaus, Fred. Cuv. whose teeth, feet, and even colours, are similar to those of the Skunk, but whose truncated muzzle resembles a Hog's snout ; the tail being reduced to a small pencil. One species only is known, the M. maliceps, Fred. Cuv., and Horsf. Java. (The Teledu). Black; the nape of the neck, a stripe along the back, and the tail white ; the dorsal stripe sometimes interrupted in the middle ; not surpassed in stench by any of the Skunks (a). Lutra, Storr. The Otters have three false molars in each jaw, a strong heel to the superior lacerator, a tubercle on the inner side of the inferior one, and a large tuberculous one above, nearly as long as it is broad. The head is compressed, and the tongue demi-asperate. They are otherwise distin- guished from all the preceding subgenera by palmated feet, and a hori- zontally flattened tail, two characters which render them aquatic. Their food is fish. L. vulgaris; Mustela lutra, L. ; Buff. VIII. xi. (The Common or Greater Otter). Brown above, whitish round the lips, on the cheeks and the whole inferior surface of the body. It is sometimes found spotted and whitish. From the rivers of Europe. Several Otters differ but little from the above. That of Carolina, * It is better figured, Hist, des Mammif. of Fred. Cuv. The Chili Skunk, Buff. Supp. VII. pi. lvii, appears to be a mere badly preserved variety of the same. See my Recherches, Sur. Ossemens Foss. IV. 469. g3T («) All these animals possess an orifice situated below the anus, which is connected with a peculiar gland: this gland secretes the unctuous matter from which the overpowering stench is exhaled; and if we are to believe in the representations of travellers, the secretions of these Skunks must be the most revolting of any fetid exhalation which nature has yet produced. Dogs are instantly stopped in their pur- suit by its emission, and if a man is so unfortunate as, when hunting them, to come in contact with the least particle of the fluid, which, when hard pressed in the chase, they are able to discharge, the garment so infected can never be used again on ac- count of the impossibility of purifying it from the horrible stench. Molina, speak- ing of the Chinge of Chili, affirms that the smell of the animals proceeds from a certain greenish oil ejected from a follicle or receptacle near the tail. — Eng. Ed. DO MAMMALIA. L. lataxina, Fr. Cuv., becomes a little larger, is sometimes more deeply coloured, and has a brownish tint beneath ; very frequently, however, there is no difference even in the shades of colour. In Brazil there are others similar in every respect to those of Carolina. That of the East Indies, the L. nair, Fr. Cuv., (the Pondicherry Otter) appears a little smoother, and is somewhat pale about the eye-brows, but it is scarcely perceptible. The Indians employ it for fishing, as we do the dog for hunting. That of Java, L. leptonyx, Horsf. (the Javanese Otter), has a whiter throat, and this whiteness ascends on the sides of the head so as to surround the eye. In that of the Cape, L. capensis, Fr. Cuv., the white on the throat, sides of the head and neck, is purer and more extended ; the end of the nose is even marked with it : what particularly distinguishes it, however, is that, at least at a certain age, it has no nails, a character on which M. Lesson has founded his genus Aonyx. Young individuals how- ever have been brought from the Cape that have nails ; it remains to be ascertained whether or not they are of the same species. Mustela Intra brasiliensis, Gm. (The American Otter). Brown or fawn-coloured ; throat white or yellowish ; a little larger than the European Otter ; the body is also longer, and the hair shorter. It is distinguished by the end of the nose, not being naked as in most animals, but being covered with hair like the rest of the chanfrin. From the rivers of both Americas. Mustela bOris, L. ; Schreb. CXXVIII*. (The Sea-Otter). Size, double that of the European species ; body much elongated ; tail one-third the length of the body ; the hind feet very short. Its blackish fur, with a marked velvety character, is the most valuable of all the furs ; it is often whitish on the head. The English and Russians go in search of this animal in the whole of the northern portion of the Pacific Ocean, for the purpose of making a traffick in its skin with China and Japan. It has no more than four incisor teeth below, but its grinders resemble those of the other Otters. The second subdivision of the Digitigrada has two fiat tuberculous teeth behind the superior lacerator, which is itself furnished with a large heel. They are carnivorous, but do not exhibit a courage proportioned to their strength, and frequently feed on carrion. The caecum is always small. Canis, Lin. Dogs have three false molars above, four below, and two tuberculous teeth behind each of the carnivori ; the first of these upper tuberculous teeth is very large. Their superior carnivorous has only a small inner tubercle, but the posterior portion of the inferior is altogether tuberculous. The tongue is soft; the fore-feet have five toes, and the hind ones four. * This figure, apparently drawn from a badly prepared specimen, presents an ex- aggerated resemblance to the Seal, a circumstance by which some naturalists have been induced to believe it should be placed near that genus — its whole organization, however, is that of the Otter. See Ev. Home, Phil. Trans. 1796. CARNARIA. i)l C. familiaris, L. (The Domestic Dog). Distinguished by his recurved tail, otherwise varying infinitely, as to size, form, colour, and quality of the hair. He is the most complete, singular, and use- ful conquest ever made by man ; the whole species has become his property ; each individual is devoted to his particular master, assumes his manners, knows and defends his possessions, and remains his true and faithful friend till death ; and all this, neither from con- straint nor want, but solely from the purest gratitude and the truest friendship. The swiftness, strength, and scent of the Dog have ren- dered him man's powerful ally against all other animals, and were even, perhaps, necessary to the establishment of society. Of all animals, he is the only one which has followed man through every region of the globe. Some naturalists think the Dog is a Wolf, and others, that he is a domesticated Jackall ; and yet, those dogs which have become wild again in desert islands resemble neither the one nor the other. The wild dogs, and those that belong to savages, such as the inhabitants of New Holland, have straight ears, which has occasioned a belief that the European races which approach the most to the original type, are the Shepherd's Dog and Wolf Dog; but the comparison of the crania indicates a closer affinity in the Mastiff and Danish Dog, subsequently to which come the Hound, the Pointer, and the Ter- rier, differing between themselves only in size and the proportions of the limbs. The Greyhound is longer and more lank, its frontal si- nuses are smaller, and its scent weaker. The Shepherd's Dog and the Wolf Dog resume the straight ears of the wild ones, but with a greater cerebral developement, which continues to increase together with the intelligence in the Barbet and the Spaniel. The Bull Dog, on the other hand, is remarkable for the shortness and strength of his jaws. The small pet-dogs, the Pugs, Spaniels, Shocks, &c, are the most degenerate productions, and exhibit the most striking marks of that power to which man subjects all nature*. The dog is born with his eyes closed ; he opens them on the tenth or twelfth day ; his teeth commence changing in the fourth month, and his full growth is attained at the expiration of the second year. The period of gestation is sixty-three days, and from six to twelve pups are produced at a birth. The dog is old at fifteen years, and seldom lives beyond twenty. His vigilance, bark, singular mode of copulation, and susceptibility of education, are well known to every one. C. lupus, L. ; Buff. VII. i. (The Wolf). A large species, with a straight tail ; legs fawn-coloured, with a black stripe on the fore- legs when adultf ; the most mischievous of all the carnaria of Eu- rope. It is found from Egypt to Lapland, and appears to have pass- ed into America. Towards the north, in winter, its fur becomes white. It attacks all our animals, yet does not exhibit a courage proportioned to its strength. It often feeds on carrion. Its habits and physical developement are closely related to those of the dog. * See Fr. Cuv. Ann. Mus. XVIII. p. 333, et seq. f This stripe is more or less strongly marked on the Jackall, Mexican Wolf, &c. 92 MAMMALIA. C. tycoon, L. ; Buff. IX, xli. (The Black Wolf). Also inha- bits Europe, and is sometimes, though rarely, found in France*. The fur is of a deep and uniform black, with a little white at the end of the muzzle, and a small spot of the same colour under the breast. It is said to be more ferocious than the common wolf. C. mexicanus, L. (The Mexican Wolf). Reddish grey, mixed with black; circumference of the muzzle, under part of the body, and the feet white ; size that of the Common Wolff. C.jubatus, Cuv. ; Agoura-Gouazou, Azzar. (The Red Wolf). A fine cinnamon-red; a short black mane along the spine. From the marshes of South America. C. aureus, L. ; Schreb. XCIV. (The Chacal or Jackal). Less than the preceding ; the muzzle more pointed ; of a greyish brown ; thighs and legs of a light fawn colour; some red on the ear; the tail scarcely reaching further than the heel. It is a voracious ani- mal, which hunts like the dog, and in its conformation, and the faci- lity with which it is tamed, resembles the latter more closely than any other wild species. Jackals are found from the Indies and the environs of the Caspian sea, as far as and in Guinea; it is not cer- tain, however, that they are all of one species. Those of Senegal, for instance, C. anthus, Fr. Cuv. Mammif., stand higher, appear to have a sharper muzzle, and the tail a little longer. Foxes may be distinguished from the Wolf and Dog by a longer and more tufted tail, by a more pointed muzzle, by pupils, which, during the day, form a vertical fissure, and by the upper incisors being less sloping. They diffuse a fetid odour, dig burrows, and attack none but the weaker animals. This subgenus is more numerous than the preceding one. C. vulpes, L. ; Buff. VII. vi. (The Common Fox). More or less red; tip of the tail white; found from Sweden to Egypt. Those of the north have merely a more brilliant fur. There is no constant difference to be observed between those of the Eastern continent and those of North America. The Calopex, Schreb. XCL, or the Collier, which has the end of the tail black, and is found in the same countries as the common one ; the Renard croise, Id. XCL A, or the Cross Fox, which is only distinguished by a streak of black along the spine and across the shoulders ; the Fox the French fur- riers call the Turk, which is of a yellowish grey, with the end of the tail white, are, perhaps, mere varieties of the common one. The following species, however, are very distinct. [See App. VII. of Am. Ed.] C. Azurce, Pr. Max ; Aguarachai, Azz. (The Brazil Fox). Grey ; * We have seen four individuals taken and killed in France. It must not be con- founded with the Black Fox, among whose synonymes Gmelin has placed it. [See Append. VI. of Am. Ed.] f This character is taken from a specimen brought from Mexico, and presented to the Cabinet du Roi by M. de Humboldt. Those which have been drawn by au- thors from the bad figure of Recchi, inserted in Hernandez, p. 479, must be reject- ed. Messrs. Say and Harlan, Faun. Amer., mention two other species of wolves, Can. latrans and Can. nubilus, which require to be examined and compared. [See Append, ut sup. of Am. Ed. CARNARIA. 9.j sides of the neck reddish ; a black line commencing on the nape of the neck, and extending along the middle of the tail. C. corsac, Gm. ; Buff. Supp. III. xvi, under the name of Adive. (The Corsac). A pale yellowish grey ; a few blackish waves at the base of the tail; tip of the tail black; jaw white. Common on the vast heaths of central Asia, from the Volga to India. It has the habits of the Fox, and never drinks. I suspect the Abouhossein of Nubia — Canis pallidus, Ruppel, pi. xi — is the same animal. There is also in the prairies of North America, a little Fox, C. velox, Har. and Say; F. Am., 91, which lives in burrows, but which appears to differ from the Corsac by the colours : a blackish tail, &c. C. cinereo-argenteus, Schreb. XCII. A. (The Tri-coloured Fox of America). Ash-coloured above ; white beneath ; a cinnamon-red band along the flanks. From all the warm and temperate parts of the two Americas. C. argentatus. (The Silver or Black Fox)*. Black; tips of the hairs white, except on the ears, shoulders, and tail, where they are of a pure black. The end of the tail is all white. From North America. Its fur is most beautiful, and very costly. C. lagopus, L. ; Schreb. XCIII. (The Blue Fox or Isatis). Deep ash-colour; the under surface of the toes hairy -j-; often white in winter. From the north of both continents, particularly from Norway and Siberia ; much esteemed for its fur. C. mesomelas^, Schreb. XCV. (The Cape Fox). Fawn-coloured on the flanks ; middle of the nose black, mixed with white, termi- nating in a point behind; the ears red as well as the feet; the two posterior thirds of the tail black, &c. The interior of Africa produces Foxes remarkable for the size of their ears, and the strength of the hairs of their mustachios ; they are the Megalotis of Illiger. There are two known, the C. megalotis, Lalande; a Cape species, something smaller than our common Fox, higher on its feet; yellowish grey above, whitish beneath; the feet, tail, and a dorsal line black. C. zerda, Gm., or Fennec of Bruce; Buff. Supp. III. xix. Ears still larger ; a small species of an almost white fawn colour, which burrows in the sands of Nubia §; its hair is woolly, and extends un- der the toes. Finally, we may place after the Dogs, as a fourth subgenus, distin- guished by the number of toes, which is four to each foot, the Hycena venatica, Bursch. ; H. picta, Temm. ; An. Gen. des Sc. Phys. III. (The Wild Dog of the Cape). It has the dental sys- * Gmel. has confounded it with the Black Wolf, under the name of Canis lycaon. ■j- Several of the Foxes, and even the common one, have hair under their feet in the north. X Gmelin has confounded it with the Adive of Button, which is a factitious species, and does not differ from the Jackal. § Bruce's figure, copied by Button, and subsequently by all his compilers, greatly exaggerates the size of the ears. We have at last a good figure and exact descrip- tion of this animal in the Voy. of Ruppel, Zoolog. pi. iii. 94 MAMMALIA. tern of the Dog and not that of the Hyena ; a long and thin form ; the fur mottled, with white and fawn colour, grey and black ; size of the Wolf, large ears with black tips, &c. It is gregarious, and fre- quently approaches Cape Town, devastating its environs. VlVERRA. The Civets have three false molars above and four below, the anterior of which sometimes fall out; two tolerably large tuberculous teeth above, one only below, and two tubercles projecting forwards on the inner side of the inferior carnivorous, the rest of that tooth being more or less tuber- culous. The tongue is bristled with sharp and rough papillae. Their claws are more or less raised as they walk, and near the anus is a pouch more or less deep, where an unctuous and frequently an odorous matter oozes from peculiar glands. They are divided into four subgenera. Viverra, Cuv. In the true Civets the deep pouch situated between the anus and the organ of generation, and divided into two sacs, is filled with an abundant pommade of a strong musky odour, secreted by glands which surround the pouch. This substance is an article of commerce, and is used by the per- fumers. It was more employed when musk and ambergris were un- known. The pupil of the eye remains round during the day, and their claws are only semi-retractile. V. civetta, L. ; Buff. IX, xxxiv. (The Civet). Ash-coloured, irregularly barred and spotted with black; the tail less than the body, black towards the end, with four or five rings near its base; two black bands encircling the throat, and one surrounding the face ; a mane along the whole length of the spine and tail that bristles up at the will of the animal. From the hottest parts of Africa. V. zibetha, L. ; Buff. IX. xxxi. (The Zibet). Ash-coloured, spotted with black; black half-rings on the whole tail; black bands on the sides of the neck ; no mane. From the East Indies. Genetta, Cuv. In the Genets the pouch is reduced to a slight depression formed by the projection of the glands, and has scarcely any visible excretion, al- though an odour is diffused from it that is very perceptible. In the light the pupil forms a vertical fissure, and the nails are completely retractile, as in the Cat. V. genetta, L. (The Common Genet). Grey, spotted with brown or black, the muzzle blackish ; white spots on the eye-brows, cheeks, and each side of the end of the nose ; tail the length of the body, annulated with black and white, the black rings being from nine to eleven in number. Found from the south of France to the Cape of Good Hope, differing in the size and number of the spots in the bands along the shoulder and neck, as well as in the lines on the nape of the neck, &c*. It frequents the edges of brooks, near springs, &c. The skin forms an important article of trade. * The best figure of a Genet is tliiit given by Pennant, Synops. No. 172, Hist. CARNARIA. 95 V. Unsaiig, Hardwick, Lin. Trans. XIII. pi. xxiv; Felis gracilis, Horsf. Java. (The Javanese Genet). Several irregular, brown, transverse bands on the body, and seven rings round the tail. V. fossa, Buff. XIII. xx. (The Fossane of Madagascar). Tail, flanks, and all above, fawn colour ; the legs and all beneath a yellow- ish white ; reddish brown spots, those on the back forming four lon- gitudinal bands ; tail semi-annulated with red, and only half the length of the body*. V. rasse, Horsf. Jav. (The Rasse). Legs brown ; body greyish brown, with small brown spots united on the crupper, and forming five longitudinal lines. Tail shorter than the body, annulated with black and white, the black rings six or seven in number f. The hair is harsher than in the preceding species. The Paradoxurus, Fr. Cuv. has the teeth and most of the characters of the Genets, with which it was a long time confounded; it is however more stout-limbed; the feet are semi-palmate, and the walk nearly plantigrade ; but what particularly dis- tinguishes it is the spiral inclination of the tail, which is not prehensile. Only one species is known, the P. typus, Fr. Cuv. (The Pougoune of India). A yellowish- brown, with some spots of a deeper brown than the rest; the feet, muzzle, and part of the tail blackish; eye-brows white, and a white spot under the eye. The French of Pondicherry call it the Palm Martin or Marte des palmier s "£ . Mangusta, Cuv. — Herpestes, Blig. The pouch voluminous and simple ; the anus pierced in its depth. The hairs are annulated with light and obscure tints, which determine their general colour on the eye. The Mangouste of Egypt, so celebrated among the ancients under the name of Ichneumon; Viverra ichneumon, L. ; Buff. Supp. III. No. 2S0, under the improper name of Fussane. It is the variety most frequently brought from the Cape. There is another taken from a young specimen, Brown, 111. pi. xliii, still under the name of Fossane. It is distinguished by its whitish and not brown legs, and we have seen a similar one from Senegal. That of Burl*. IX. xxxvi, has not the bands on the neck and shoulders sufficiently marked. The num- ber of black rings on the tail varies from nine to eleven. The Civette de Malacca of Sonnerat, Voy. II. pi. xxxix, which is the same as the Genette du Cap, Buff. Supp. VII. pi. lviii, and the Chat bisaam of Vosmaer, of which Gmelin has made as many species, appear to be common Genets. * Description taken from the original sent to Buffon by Poivre, and engraved, Hist. Nat. XIII. pi. xx. The description of Daubenton is correct so far as regards the distribution of the spots; but he calls them black, whereas they are reddish. Be- sides, this animal can hardly be the fossa of Flacourt, which that author states is the size of the Badger. The Fossane has the same furrow as the Genet, notwithstand- ing the assertion of Poivre to the contrary. f It is probably V animal du muse of La Peyronie, Acad, des Sc. 1728, pi. xxiv. p. 464, which had been confounded with the Zibeth — but that animal is larger, and has other colours. To this division we must refer the Viv. fasciata, Gm.; Buff. Supp. VII. lvii. X It is the pretended Genette de France of Buffon, Supp. IF I. pi. xlvii, the Civette a bandeau of Geoff. '.)(', MAMM VLIA. xxvi. is frrf-y, with a long tail, terminated with a black tuft; it is larger than our cat, and idei : > a marten. It chiefly hunts for the eggi of the crocodile, but also feeds on all sorts of small ani- mal:;; brought up in bouses, it hunts mire, reptiles, &c. By the Europeans al Cairo il i called Pharaoh's Rat; by the natives, Nems. The antieht tradition of its jumping down the throat of the croco- dile, to dei troy it, r entirely fabulout ■ The Mangouete of India; Viv. mmgos, Lin.; Buff. XIII, xix; ami that of the Cape, Viv. cafra, Gbn.; Schreb. CXVI. B., are smaller, both having a pointed tail, and a grey or brown fur, the lat- ter being more of an ashy, and the former more of a fawn colour, having, besides, some red about the cheeks and jaws. The Mangouste of India is celebrated lor its combats with the mosl dangerou serpents, and lor having led us to the knowledge of the Ophiorhiza rnongo , a an antidote to their poison. There i :| l o the Mangouste of Java H. Javanicus, reddish brown; cheeks of a che tnut-red; throat more fawn-coloured: a large one, from the marshes of the Cape //. pahidinosus, of an al- ino i iiinl'onn reddish-brown, verging to a black, a little lighter on the chin: a third.from the Cape //. penicillatus, of a greyish lawn (•(dour, tip of the tail white: one from Senegal //. albicaudus, grey, tail all white: it is difficult, however, to establish very specific difference bi tween thei e animals. I{ vz.i/.a, HUg. The Surikates have a strong K emblance to the Mangoustes, even to the tints and transverse streaks of the hair, bu1 are distinguished from them, and from all the Carnivora of which we have hitherto spoken, by having onlj foui toes to each foot. They also are higher on their logs, and they have not the small molar immediately behind the canine tooth. Their pouch extends into the anus. One pecies only is known, a native of Africa — Viv. tetradactyla, (Jin. ; Hull'. X 1 1 J, viii, a little less than the Mangouste of India*'. Gbobsaechus, Fred. Cuv. The muzzle, teeth, pouch, and walk of the Surikates, the Iocs and ge- nital organs of the Mangoustes. One species only is known Crossavchus obscuru8 f Fred. Cuv., from Sierra laone, of the size of the Surikale; greyish brown ; cheeks a little paler, and a hairy tale. We should here mention a singular animal from the south of Africa, known only while young; which, to the live anterior toes, and the lour hind one:;, and the slightly elongated head of the ci- vets, adds the raised feet, the short hind ones, and the mane of the hyena,; it. also singularly resembles the striped hyena in the colour, of its fur. The thumb of the fore fool, is short, and higher; it is the * The Zenil of Sonnerat, Voy. II, pi. xcii, appears to differ from the Surikatc, merely becaui i it ia roughly drawn. CARNARIA. 9 Protelcs Lalandii, Isid. Geoff. Mem. du Mus. XI. 354, pi. xx. In- liabits caverns. The individual specimens that have been examined, and which were all young, had but three small false molars, and one small tu- berculous posterior molar. It seems as though their teeth had never come to perfection, as often happens in the Genets*. * See my Ossemens Fossiles, torn. IV. p. 388. ffcW (a) In this second subdivision of the digitigrade animals, which is here con- cluded, are found many species particularly recommended to our attention. As a vast collection of dogs of different countries are now in the two Zoological Gardens, it may he convenient to our readers to give the simple classification of these interesting animals, as it has been recently established by M. F. Cuvier. He forms the whole species of dogs into three groups : — 1. The Matins, characterized as follows — head elongated, sides of the cranium approaching each other in the anterior direction, and the condyles of the inferior jaw horizontally directed with respect to the position of the teeth in the upper jaw. The specimens of this group are — the New Holland Dog, the French Matin, the Danish Dog, the Greyhound, including all the varieties of the latter, and the Albanian Dog. 2. Spaniels, in which the head is elongated, but not to the same extent as in the Matins, nor do the side bones of the head approximate each other, but they sepa- rate more widely, swelling out in such a way as to increase the anterior part of the cavity of the cranium, a circumstance that would admit of the supposition that these animals owe their superior intelligence to the greater developement of the hemis- pheres of the brain. This group includes the various spaniels, such as the King Charles's breed, the Water Spaniel, the Hound, Bloodhound, Foxhound, Harrier, and the Beagle, which is a particular breed of the Harrier, the Pointer, Turnspit, Shepherd's Dog, Wolf, Siberian and Esquimaux Dogs, the Alpine and Newfoundland Dog, the Setter, Terrier, and the Alco. 3. Dogues. In this third group the muzzle is shortened, the cranium is very high, and smaller than in other dogs, and the sinuses between the walls of the frontal bone vary considerably. It includes the Bull Dog, Mastiff, Pug Dog, Iceland Dog, Little Danish, Bastard Pug, the Artois, the Barbary, and Dog of Andalusia. Some Esquimaux, Australian, and other varieties of foreign Dogs, may be seen in both the Zoological Gardens. The Wolf Species, belonging to this subdivision, are found in various parts of the continent of Europe at present, but they offer this remarkable peculiarity m their history, that their race, which was once exceedingly numerous and formidable in these countries, has been wholly extirpated from them. The last native wolf which is re- corded to have been seen here, was in 1710, in Ireland. A male and female wolf, the one a native of France, the other of Russia, are in the Zoological Gardens. The property possessed by Civet Cats, the genus Viverra of this subdivision, of secreting an odoriferous substance, which was once used in medicine, but is now ex- clusively employed in perfumery, merits some allusion. A deep bag, situated be- tween the anus and the organs of generation, is divided into two cavities, into which two glands secrete the peculiar matter called civet. The substance is a thick, unctuous matter, with an odour very much resembling that of amber. In the fresh state it is white, but, after some time, becomes yellow, and acquires a very agreeable odour. Chemical investigation has shewn that the odour depends on a volatile oil mixed with some other ingredients, from which it may be separated by distillation in water. The oil, when thus isolated, is of a clear yellow colour, has the strong odour of civet, together with an acid and burning taste. We may add, inthis place, that the secretion from the Skunks (see p. 88), has been found, on analysis, to con- sist of two oils, which may be separated from each other; the one is an oil resem- bling the amber in colour", giving out a most revolting garlic smell, so that in the .smallest possible quantity it is perfectly insupportable, and communicates its stench even to water with which it is mixed. The thick oil, which is another of its ingre- dients, has no flavour whatever. — Eng. En. VOL. I. H 98 MAMMALIA. The last subdivision of the Digitigrada has no small teeth of any kind behind the large molar of the lower jaw. The animals contained in it are the most cruel and sanguinary of the class. They form two genera. Hyzena, Storr. The Hyenas have three false molars above and four below, all conical, blunt, and singularly large; their superior carnivorous tooth has a small tubercle within and in front, but the inferior has none, presenting only two stout trenchant points: with these powerful arms they are enabled to crush the bones of the largest prey. The tongue is rough ; each foot, has four toes like that of the Surikate ; and under the anus is a deep and glandular pouch, which induced some of the ancients to consider them as hermaphro- dites. So powerful are the muscles of the neck and jaw, that it is almost impossible to wrest any thing from between their teeth that they have once seized, and, among the Arabs, their name is the symbol of obstinacy. It sometimes happens that an anchylosis of the cervical vertebrae is the consequence of these violent efforts, and this has caused it to be said that they have only one single bone in the neck. They are nocturnal animals, inhabiting caves ; are extremely voracious, and feed chiefly on dead bodies, which they seek for even in the grave. A thousand superstitious tradi- tions are connected with them. Three species are known, the H. vulgaris, Buff. Supp. III. xlvi. (The Striped Hyena). Grey; blackish or brown stripes crosswise; a mane along the whole of the nape of the neck, and black, that stands erect when the animal is angry. It is found from India to Abyssinia and Senegal. H. hrunnea, Thumb., Acad, of Stockh. 1820, part I. pi. ii; H. villosa, Smith. Lin. Trans. XV. pi. xix. (The Brown Hyena). Of a deep greyish brown ; black stripes on the legs only. From the south of Africa, where the inhabitants of the Cape call it le Loup du rivage, or the Shore Wolf. H. croeuta, Schreb. XCVI. B. (The Spotted Hyena). Grey or reddish, sprinkled with black spots. It is likewise from the south of Africa, and is the Tiger Wolf of the Cape. There have lately been found in several caverns of France, Ger- many, and England, many bones of a lost species of Hyena — H. spelcea, which appears to have resided there, and to have left the bones of many other animals, which bear evident marks of its teeth, and even its own fasces*. * See Buckland, Reliquiae Diluvianae, and Vol. IV. of my Oss. Foss. 2d ed. (jCff* (a) The learned professor mentioned in the foregoing note, discovered, in 1822, in Kirkdale cave, Yorkshire, beneath a muddy surface, a great quantity of the bones of different animals, a remarkable proportion of which belonged to Hyaenas. The conclusions to which this discovery led were as follows : — that this cavern was, before the deluge, the retreat of Hyaenas, the species of which have been long ex- tinct; that the teeth and fragments of the bones with which the remains of the Hyae- nas were found blended, belonged to the elephant, rhinoceros, hippopotamus, horse, ox, deer, fox, water-rat, and several birds, all these animals having been dragged into the cave to be devoured by the Hyaenas. At least, this motive seems to have been strongly indicated by the fact, that the fragments shewed proofs of having been CARNARIA. Ji!) Felis, Linn. Of all the Carnaria the Cats are the most completely and powerfully armed. Their short and round muzzle, short jaws, and particularly their retractile nails, which, being raised perpendicularly, and hidden between the toes, when at rest, by the action of elastic ligament, lose neither point nor edge, render them most formidable animals, the larger species espe- cially. They have two false molars above, and two below: their superior carnivorous tooth has three lobes, and a blunted heel on the inner side, the inferior, two pointed and trenchant lobes, without any heel : they have but a very small tuberculous tooth above, without any thing to correspond to it below. The species of this genus are very numerous and various with regard to size and colour, though they are all similar with respect to form. We can only subdivide them by referring to the difference of size and the length of the hair, characters of but little importance. At the head of the genus we find F. leo, L.; Buff. VIII. i. 11. (The Lion). Distinguished by its uniform tawny colour, the tuft of hair at the end of the tail, and the flowing mane which clothes the head, neck, and shoulders of the male. Of all beasts of prey, this is the strongest and most coura- geous. Formerly scattered through the three parts of the old world, it seems at present to be confined to Africa and some of the neigh - bouring parts of Asia. The head of the Lion is more square than that of the following species. Tigers are large, short-haired species, most commonly marked with vivid spots. F. tigris, Buff. VIII. ix. (The Royal Tiger). As large as the Lion, but the body is longer, and the head rounder ; of a lively fawn colour above; a pure white below, irregularly crossed with black stripes; the most cruel of all quadrupeds, and the scourge of the East Indies. Such are his strength and the velocity of his move- ments, that during the march of armies he has been seen to seize a soldier while on horseback, and bear him to the depths of the forest, without affording a possibility of rescue. F. on^a, L. ; Azzar. pi. ix; Fred. Cuv. Mammif. (The Jaguar). Nearly the size of the Royal Tiger, and almost as dangerous ; a lively gnawed, and even fractured by particular teeth. Amongst the remains were teeth and excrements of the Hyaenas also, the existence of which has been explained on the principle that it is the ascertained habit of Hyaenas to devour the dead bodies of their own species, being, like wolves, gregarious, and hunting mostly in packs. Similar fossil remains of supposed antediluvian Hyaenas have been found in France and Germany, in caves; but the circumstances under which they have been disco- vered lead to the conclusion that the bones either belong to animals that had fallen through fissures opening into these caves, or were carried by water through subter- ranean canals. The species, which is unknown as it existed previously to the deluge, is called H. Spela?a, Cave Hyeena, and by means of that exact knowledge of the laws of animal organization which he so eminently possessed, Cuvier has been able to build up afresh the whole of the structure of this species, and has given the following description of the unseen animal in another of his great works : — size larger by a third proportion than the Hyaena rayee, Canis hycenus; the muzzle, however, is shorter, and the teeth must have been much larger, from the appearance of their fragments, which consist but of stumps, than those of the existing races. — Eng. Ed. h2 100 MAMMALIA. fawn colour above ; the flank longitudinally marked with four rows of ocellated spots, that is, with rings more or less complete, having a black point in the middle ; white beneath, transversely striped with black. Sometimes individual specimens are found black, whose rings, of a deeper hue, are only perceptible in a particular light. F. pardus, L. ; the Partialis of the ancients ; Cuv. Menag. du Mus. 8vo. I. p. 212. (The Panther). Fawn coloured above ; white beneath ; with six or seven rows of black spots, resembling roses, that is, formed by the assemblage of five or six simple spots on each flank ; the tail is the length of the body, minus that of the head. This species is scattered throughout all Africa, the southern parts of Asia, and the Indian Archipelago. In some of them the ground of the fur is black, with spots of a deeper black — F. inelas, Per., but they are not a distinct species. We have frequently seen black and fawn-coloured young ones suckled by the same mother*. F. leopardus, L. (The Leopard). From Africa; similar to the Panther, but has ten rows of smaller spots j. These two species are smaller than the Jaguar. Travellers and furriers designate them indiscriminately by the names of Leopard, Panther, African Tiger, &c.J. There is a third, peculiar to the distant parts of the East Indies, that is a little lower; tail equal in length to the body and head; spots smaller and more numerous ; the F. chalybeata, Herm. ; Schreb. CI.§. F. discolor, L. ; Buff. VIII. xix. (The Couguar or Puma). Red, with small spots of a slightly deeper red which are not easily perceived. From both Americas, where it preys on Deer, Sheep, &c. ||. Among the inferior species we should distinguish the Lynxes, * Temminck calls this species Felis leopardus. •)• The same naturalist considers our Leopard as a variety of our Panther, and confounds them under his Felis leopardus. % Buffon has mistaken the Jaguar, which he took for the Panther of the eastern continent, and has not well distinguished the Panther and the Leopard, and for this reason we cannot positively quote his pi. xi, xii, xiii and xiv. of Vol. VIII. § It is to this species that Temminck affixes the name of Panther, because he thinks Linnaeus alluded to it, when speaking of his Felis pardus in the " cauda elon- ga/a." There is one thing very certain, and that is, that the Panther, so well known to the ancients, and which was so often produced at the Roman games, could not possibly have been an animal from the extreme parts of oriental Asia. The Ounce of Buff. IX. pi. xiii (Felis uncia, Gm.), differs from the Panthers and Leopards by the inequality of the spots, which are more irregularly distributed, and partly crenate or annulated, &c. It appears to be found in Persia. We only know it by the figure of Buffon, and that which Mr. Hamilton Smith has inserted in the work of Griffith, taken from a specimen that was living in London. [| That this animal, our common Panther, does not always confine itself to sheep, &C is well known, and has lately been proved, January, 1830, by an unprovoked attack upon an unfortunate woman in Pennsylvania. The ferocious brute seized upon her as she was passing along the road, and killed her in an instant. See Griff., part V. p. 4,18— An American trsnslation. CARNARJA. 101 which are remarkable for the pencils of hair which ornament their ears. Four or five different kinds of them are known in commerce by the name of Loups Cerviers, which have long been confounded by naturalists (Felis lynx, L.)., and whose specific limits are even not yet perhaps well ascertained. They all have a very short tail, and a skin more or less spotted. The most beautiful, which are as large as a wolf — F. cervaria, Temm., come from Asia by the way of Russia, and have a slightly reddish-grey fur, finely spotted with black. Others from Canada and the north of Sweden — F. borealis, Temm., have the fur very much tufted, extending even under the feet ; of an ash-coloured grey, and with scarcely any spots. The Lynx of the temperate parts of Europe— F. lynx, Temm., which has almost disappeared from its populous districts, but which is still found in the Pyrenees, in the mountains of Naples, and, as it is said, even in Africa ; has a red fur, spotted with brown. In these three species or varieties, the end of the tail is black. It is thought there is a lynx of the south of Europe — Fells pardina, Oken, which may be considered distinct. It is smaller, not so hairy, fur red, mottled with black, and the tail spotted like the body. We find also in North America, the F. rufa, Guld'. Schreb. CIX. B; F. montana, L. (The Bay Lynx.) A reddish fawn or greyish colour, mottled with brown; brown waves on the thighs ; tail annulated with black or brown ; ra- ther smaller than the Lynx*. F. chaus, Guld. ; Schreb. CX. (The Chaus, or Lynx of the Marshes). Is of a yellowish grey-brown; the hind part of each leg blackish; tail reaches to the hamstrings, and is annulated at the ex- tremity with black. Inhabits the Caucasian marshes, those of Persia and of Egypt, pursues birds, &c. It is now thought we should separate from the above species the Booted Lynx — F. caligata, Temm., Bruce, pi. xxx, which is some- what smaller, and has a little longer tail ; the external surface of its ears is red. It is, at least, a closely allied species, and has the same habits. F. caracal, L. ; Buff. IX. xxiv., and Supp. III. xlv. (The Ca- racal). Of an almost uniform vinous red. From Persia, Turkey, &c. It is the true Lynx of the ancients. The inferior species, which are deprived of the pencils on the ears, are more or less similar to our common cat; such are F.pardalis, L. ; Buff. XIII. pi. xxxv. and xxxvi. (The Ocelot). Rather lower on its legs than most of the others ; grey, with large * M. Rafinisque also indicates a Lynx fasciatus, a L. aureus, a L.floridanus, a L. montanus, and M. Temminck a Felis aurata, which must all belong to this, little tribe. \(Y> MAMMALIA. fawn coloured spots bordered with black, forming oblique bands on the flank. From America. F. mitisy Fr. Cuv. (The Chati). Marked with unconnected, triangular, fawn coloured spots, edged with black. F. eafra. (The Cat of Caffraria). Stands high on its legs ; grey, transversely striped with black. F. serval, Buff. XIII. xxxv. (The Serval). Yellowish, with ir- regular black spots. From Africa. F. jaguarondi, Azzara, Voy., pi. 9. (The Jaguarondi). Body long; and altogether of a blackish brown. From the forests of South America. F. catus, L. ; Buff. VI. i. et seq. (The Domestic Cat). Is ori- ginally from the forests of Europe. In its wild state it is of a grey- ish brown, with darker transverse undulations; below pale; the in- sides of the thighs and of all the feet, yellowish; three bands on the tail, its inferior third blackish. In a domestic state it varies, as is well known, in colours, in the length and fineness of the hair, but infinitely less so than the dog; it is also much less submissive and affectionate*. We might also place in a separate subgenus, a species whose head is rounder and shorter, and whose nails are not retractile, the Felis jubata, Schreb. 105, and better, Fel. guttata, Id. 105, b, (The Hunt- ing Leopard), which is the size of the Leopard, but longer bodied, and stands higher ; the tail long, annulated at the end ; the fur fawn colour, mottled with small uniform black spots, a black streak reach- ing from the eye to the angle of the mouth. The disposition of this animal differs from that of the remainder of the genus in being ex- tremely mild and docile. The AMPHIBIA Will form the third and last of the small tribes into which we divide the Carnivora. Their feet are so short and so enveloped in the skin, that the only service they can render them on land, is to enable them to crawl; but as the intervals of the fingers are occupied by membranes, they are excellent oars ; and, in fact, these animals pass the greater portion of their time in the water ; never landing, except for the purpose of basking in the sun, and suckling their young. Their elongated body; their very move- able spine, which is provided with muscles that strongly flex it; their narrow pelvis ; their short hair, that adheres closely to the skin, all unite * The species, more or less allied to the cat, are very numerous in the two conti- nents; but all those that are given in catalogues are very far from being authentic, and sufficiently distinguished from each other. We may, however, consider as such, those of which we have good figures. The Margay, Buff. ; Felis tigrina, Gm., Buff. XIII.; Schreb. 10(i. — Fel.macroura, Pr. Max., Brazil, pi. xi. — Felis sumatrana, Horsl. — Fel. javanensis, Id. — Fel. torquata, Fred. Cuv. — Fel. colocolo, Fred. Cuv. Maminif., CARNARIA. 103 to render them good swimmers; and all the details of their anatomy con- firm these first indicia. We have as yet distinguished two genera only, Phoca and Trichechus. Phoca, Lin. Seals have six or four incisors above, four or two below, pointed canini and grinders to the number of twenty, twenty-two, or twenty-four, all trenchant or conical, and without any tuberculous part whatever; five toes to all the feet, the anterior ones regularly decreasing in length from the thumb to the little toe, while in the hinder feet the thumb and the little toe are the longest, and the intermediate ones the shortest. The fore feet are enveloped in the skin of the body as far as the tarsus, the hinder ones almost to the heel. Between the latter is a short tail. The head of a seal bears a resemblance to that of a dog, whose intelligence and soft ex- pressive look it also possesses. It is easily tamed, and soon becomes at- tached to its keeper, or those who feed it. The tongue is smooth, and sloped at the end, the stomach simple, caecum short, and the intestinal canal long, and tolerably regular. These animals live on fish ; always eat in the water, and close their nostrils when they dive by a kind of valve. As they remain a long time under water, it was supposed that the foramen ovale remained open, as in the human foetus — but it is not so : there is, however, a large venous sinus in the liver, which must assist them in diving, by rendering respiration less necessary to the motion of the blood. Their blood is very abundant and very black. Phoca, properly so called, or, without external ears. The true Phocae have pointed incisors ; all the toes enjoy a certain de- gree of motion, and are terminated by pointed nails planted on the edge of the membrane, which unites them. They are subdivided from the number of their incisors. The Caloce- phala, Fr. Cuv., have six above and four below; such is the Phoea vitulina, L. ; Buff. XIII. xlv., and Supp. VI. xlvi; Ph. littorea, Thienem. pi. vi. (The Common Seal). From three to five feet in length ; of a yellowish grey, more or less shaded and spotted with brown, according to its age ; sometimes brownish, with small yellow spots. When very old it becomes whitish. Common on the coast of Europe in great herds. It is also found far to the north ; we are even assured that it is this species which inhabits the Caspian sea, and the great fresh water lakes of Russia and Siberia, but this assertion does not appear to be founded on an exact comparison. In fact, the European seas contain several Phocae, which have long been confounded, some of which are perhaps mere varieties of the others. Thus, some of them have the back covered with small clouded, confluent, brownish spots, on a yellowish ground — Ph. hispida, Schreb. 80*. These are the most common ones of the northern ocean. In others again the ground is dark, traversed with undulat- * I suspect we should refer to it the Ph. sropulico/a, Thienem, pi. v. J 04 MAMMALIA. iiig lines, which sometimes forms rings — Ph. annellata, Nils., Thie- nem., pi. ix — xii; Ph.foetida, Fabr*., &c. A species more easily recognised is the Ph. groenlandica, and P. oceanica; Eged. Groenl. fig. A, p. 62; Lepechin, Act. Petrop. I, part I. pi. vi — vii. ; Thieneman, pi. xiv — xxi. (The Harp Seal). Yellowish grey, spotted with brown when young, afterwards marked by an oblique black or brown scarf on each flank ; the head of the old male is black ; length five feet. From the whole north of the globe. Ph. barbata, Fabr.; Thienem., pi. i — iv. (The Bearded Seal). From the north, and surpasses all the preceding ones in its size, which is from seven to eight feet : it is grey ; browner above, with a longitudinal blackish line that forms a sort of cross upon the chanfrin. Its mustachios are thicker and stronger than the others. Ph. leueopla, Thienem., pi. xiii. (The White-nailed Seal). Is of a yellowish grey. Ph. lagura, Cuv. (The Hare-tailed Seal). Has the tail white and woolly, &c.y. Stenorhincus, Fred. Cuv. Four incisors above, and four below, the molars deeply notched into three points. One species only is known, and that is from the Austral seas — Ph. leptonix, Blain. Size of the barbata; greyish above; yellowish beneath ; nails small. Pelagus, Fred. Cuv. Four incisors also, above and below, but their grinders are obtuse cones, with a slightly marked heel before and behind. There is one of them in the Mediterranean. Ph. monachus Gm. ; Buff. Supp. VL pi. xiiij. (The Monk). From ten to twelve feet in length, of a blackish brown, with a white belly. It is particularly found among the Grecian and Adriatic Islands, and is, most probably, the species best known to the antients. Stemmatopus, Fred. Cuv. Four superior incisors, and two inferior; grinders compressed, slightly trilobate, supported by thick roots. Such is the Ph. cristata, Gm. ; Phoca leonina, Fabr. ; Eged. Groenl., pi. vi. ; Dekay, New York Lye. I, pi. vii. (The Hooded Seal). Seven or eight feet long; a piece of loose skin on the head, which can be in- flated at the pleasure of the animal, and is drawn over the eyes when * It is one of those represented by Fr. Cuv. under the name of " Phoque commun." t I only wish to mention those species which I consider sufficiently ascertained. The long catalogues of the Phocse, recently published, seem to me to multiply them a great deal too much. X It is the same individual described by Hermann, Soc. des Nat. de Berl. IV. xii, xiii. under the name of monarchus. CARNAR1A. 105 it is menaced, at which times the nostrils also are inflated like blad- ders. From the arctic ocean*. Finally, the Macrorhinus, Fr. Cuv., has the incisors of the preceding, obtuse conical molars, and the muzzle resembling a short moveable pro- boscis or snout. The largest seal known is of this subgenus ; the Ph. leonina, L. ; Sea Lion of Anson ; Sea Wolf of Pernetty, &c. Peron's Voy. I. xxxii. (The Elephant Seal) (a). From twenty to twenty-five feet in length; brown, the muzzle of the male terminated by a wrinkled snout, which becomes inflated when the animal is angry. It is common in the southern latitudes of the Pacific Ocean, at the Terra-del-Fuego, New Zealand, Chili, &c. It constitutes an important object of the fisheries, on account of the oil in which it abounds. The Otaries, Peron. Seals with external ears Are worthy of being formed into a separate genus ; because, independently of the projecting external ears, the four superior middle incisors have a double cutting edge, a circumstance hitherto unknown in any animal ; the external ones are simple and smaller, and the four inferior bifurcated. All the molars are simply conical, and the toes of the fore feet almost immov- able ; the membrane of the hind feet is lengthened out into a slip beyond each toe ; all the nails are flat and slender. Ph.jubata, Gm. ; Sea Lion of Steller, Pernetty, &c. ; Buff. Supp. VII. xlviii. From fifteen to twenty feet, and more, in length; fawn coloured; the neck of the male covered with hairs that are more frizzled and thickly set than those on the rest of the body. It might be said to be found in all the Pacific Ocean, were it not that those from the straits of Magellan seem to differ from such as are taken at the Aleutian islands. Ph. ursina, Gm. ; Buff. Supp. VII. xlvii. (The Sea Bear). Eight feet long, no mane, varying from brown to whitish. From the north of the Pacific Ocean. Other seals are found in that sea which only differ from the ursina in size and colour: such is the Petit phoqur noir of Buffon (Ph. pusilla), Buff. XIII. liii; the Yellow Seal of Shaw, &c. * The mechanism by which this inflation is effected is not yet well understood. See Dekay and Ludlow, Annals of the New York Lyceum, Vol. I. pp. 94 and 99. @3T {a) A much more full and interesting account of the Sea Elephant, under the title of Phoca proboscidea, is given by two recent French travellers, Peron and Le Sueur. This is the species of seal which forms the great material of the English seal fishers off the islands in the neighbourhood of New South Wales. The fishery is now carried on periodically, and its object is to obtain the Sea Elephant, not on account of its flesh, but for the skin and oil which it is capable of yielding. The flesh is insipid and black, but still is consumed by the natives; the tongue alone is preserved by the English seamen; for, when properly cured, it is sold as a precious luxury. The fresh blubber of this animal is in the highest esteem amongst the sailors, as an easy, speedy, and most successful local remedy in all sorts of wounds. The travellers just mentioned were informed by the Englishmen engaged in this oc- cupation at the island of King, that the animal, as soon as it has been killed, is skinned and sliced into small cube-shaped pieces, which are boiled in cauldrons ar- l()(j MAMMALIA. Trichechus, Lin.* The Morse resembles the Seal in its limbs, and the general form of the body, but differs widely from it in the teeth and head. There are no in- cisors nor canini in the lower jaw, which is compressed anteriorly to pass between two enormous canini or tusks, which issue from the upper one, and which project downwards, being sometimes two feet long, and of a proportionable thickness. The enormous size of the alveoli, requisite for holding such tremendous canini, raises up the whole front of the upper jaw, giving it the shape of a huge inflated jowl, the nostrils looking upwards, and not terminating the muzzle. The molars are all short, obliquely truncated cylinders ; there are four of them on each side, above and be- low, but, at a particular age, two of the upper ones fall out. Between the canini are two incisors, similar to the molars, which most authors have not recognised as such, although they are implanted in the intermaxillary bone. Between these again, in the young animal, are two more small pointed ones. The stomach and intestines of the Morse are very similar to those of the seal. It appears that the fucus constitutes part of its food, along with animal matters. One species only is as yet ascertained, the Trich. rosrnarus, L"j~. ; Buff. XIII. liv. ; and better, Cook, Voy. III. (The Sea Cow). It inhabits the Arctic seas, surpasses the largest ox in size, attains the length of twenty feet, and is covered with a short yellowish hair. It is sought for on account of its oil and tusks ; the ivory of which, although rough grained, is employed in the arts. The skin makes excellent coach braces J. ORDER IV. MARSUPIALIA. So many are the singularities in the economy of the Marsupialia, or pouched animals, as they are termed, which we formerly placed at the end * Trichechus, from Trix (hair), a name invented by Artedi for the sea cow. f Shaw, however, suspects, that there may be two, distinguished by the greater or less size of their trunks, and by their being more or less convergent. % Previous to my arrangement, the Lamantins and Dugongs, much more nearly allied to the Cetacea, were very properly united with the Morses. ranged along the shores, and the oil is afterwards put into casks. The Frenchmen state that the oil which they saw prepared by the English sailors was clear, and free from that rancid odour which never cau be removed almost from whale and fish oil, and that it appeared to them particularly useful as a lamp oil, in consequence of its sending forth no disagreeable smell, and also because a given quantity fed the lamp for a longer period than the same amount of other oils used for the same pur- pose. The Sea Elephants feed on cuttle fish and sea weed, and stones and gravel are found in their stomachs; very commonly, too, huge calcareous concretions, which, when seen by those who open them, excite astonishment as to the possibility of their being contained within a cavity apparently so small. — Eng. Ed. MARSUPIALIA. 107 of the Carnaria as a fourtli family of that great order, that it appears to us they should form a separate and distinct one, particularly as we ohserve in them a kind of representation of three very different orders. The first of all their peculiarities is the premature production of their young, whose state of development at birth is scarcely comparable to that of an ordinary foetus a few days after conception. Incapable of motion, and hardly exhibiting the germs of limbs and other external organs, these diminutive beings attach themselves to the mammae of the mother, and re- main fixed there until they have acquired a degree of development simi- lar to that in which other animals are born. The skin of the abdomen is almost always so arranged about the mammae as to form a pouch in which these imperfect little animals are preserved as in a second uterus ; and to which, long after they can walk, they always fly for shelter at the approach of danger. Two particular bones attached to the pubis, and interposed between the muscles of the abdomen, support the pouch. These bones are also found in the male, and even in those species in which the fold that forms the pouch is scarcely visible. The matrix of the animals of this family does not open by a single ori- fice into the extreme end of the vagina, but communicates with this canal by two lateral tubes resembling handles. The premature birth of the young appears to depend upon this singular organization. The scrotum of the male, contrary to what obtains in other quadrupeds, hangs before the penis, which, when at rest, is directed backwards. Another peculiarity of the Marsupialia is, that notwithstanding a general resemblance of the species to each other, so striking that for a long time they were considered as one genus, they differ so much in the teeth, the organs of digestion, and the feet, that if we rigorously adhered to these characters, we should be compelled to separate them into several orders. They carry us, by insensible gradations, from the Carnaria to the Roden- tia, and there are even some animals which have the pelvis furnished with similar bones ; but which, from the want of incisors, or of all kinds of teeth, have been approximated to the Edentata, where, in fact, we shall leave them, under the name of Monotremata. In a word, we would say that the Marsupialia form a distinct class, pa- rallel to that of quadrupeds, and divisible into similar orders : so that if we were to arrange these two classes into two columns; the Sarigues, the Dasyuri, and the Perameles would be opposite to the insectivorous Carnaria with long canini, such as the Tenrecs and the Moles ; the Pha- langers and the Potoroos, opposite to the Hedgehogs and Shrews; the Kanguroo, properly so called, cannot be compared with any thing; but the Phascoloniys should be opposite to the Rodentia. Finally, if we were to consider the bones of the pouch only, and regard as Marsupialia 108 MAMMALIA. all the animals that possess them, the Ornithorinci and the Echidncc would form a group parallel to that of the Edentata. Linnaeus arranged all the species he was acquainted with under his ge- nus Didelphis, a word signifying double uterus. The pouch in some re- spects is in fact a second one. The first subdivision of the Marsupialia is marked by long canini, and small incisors in both jaws, back molars bristled with points, and all the characters in general of the insectivorous Carnaria ; the animals that com- pose it are also perfectly similar to the latter in their regimen. Didelphis, Lin. The Opossums*, which, of all the Marsupialia, have been the longest known, form a genus peculiar to America. They have ten incisors above, the middle ones being rather the longest, and eight below; three anterior compressed grinders and four posterior bristled grinders, the superior ones triangular, and the inferior oblong, which, with the four canini, make in all fifty teeth, the greatest number hitherto observed in quadrupeds. Their tongue is papillated, and their tail prehensile and partly naked. Their hinder thumb is long, and very opposable to the other four toes, from which circumstance these animals are sometimes styled Pedimana; they have no nail. Their extremely wide mouth, and great naked ears, give give them a very peculiar physiognomy. The glans penis is bifurcated. They are fetid and nocturnal animals, whose gait is slow ; they remain on trees, and there pursue birds, insects, &c, though not despising fruit. Their stomach is simple and small, their caecum of a middling size, and without any enlargements. The females of certain species have a deep pouch in which are the mamma?, and in which they can enclose their young. Did. virginiana, Penn. Hist. Quadr. 302 -f. (The Opossum). Almost the size of a cat ; fur, a mixture of black and white ; ears, one side black, and the other white ; head nearly ail white. Inhabits all America ; steals at night into villages ; attacks fowls, eats their eggs, &c. The young ones at birth, sometimes sixteen in number, weigh only a grain each. Although blind and nearly shapeless, they find the mammae by instinct, and adhere to them until they have at- tained the size of a mouse, which happens about the fiftieth day, at which epoch they open their eyes. They continue to return to the pouch till they are as large as rats. The term of gestation in the uterus is but twenty-six days*. Did. Azzarce, Teram. (The Gamba, or the Great Opossum of of Paraguay and Brazil). Differs from the preceding in the black * Carigueia, according to Marcgrave, is their Brazilian name, whence we have Sariguoi, Cerigon, Sarigue. They are called Micoure in Paraguay; Manicou in the islands; Opossuvi in the United States; Thlaqualzin in Mexico. f It is the Sarigue cles Illinois, and the Sarigue a longs poils; Buff. Supp. VII. p. xxxiii and xxxiv; Did. marsupialis, Schreh. pi. clxv. * See the letter of Dr. B. S. Barton to M. ltoume on the gestation of the Opossum. MARSUP1ALIA. 109 which marks the muzzle and nearly the whole of the ears ; the tail is also longer. Did. marsupialis, and Did. cancrivora, L. ; Buff. Supp. III. liv. (The Crab-eating Opossum). Size of the preceding; yellowish, mixed with brown, with brown hairs ; a brown streak on the chanfrin. It frequents the marshes of the sea coast, where it feeds chiefly on crabs*. Did. opossum, Z. ; Buff. X. xlv, vlvi. (The Four-eyed Opossum). Chestnut above, white below, a white or pale yellow spot over each eye ; posterior third of the tail white ; larger than a large rat. Other species possess no pouch, having a mere vestige of it in a fold of the skin on each side of the abdomen. They usually carry their young on their backs, the tails of the latter being entwined around that of the mother. Did. nudicauda, Geoff. ; D. myosuros, Temm. (The Bare-tailed Opossum). Fawn-coloured: tail very long, and naked even at its base; two whitish spots over each eye, one beneath. Did. cayopollin\, Did. philander, and Did. dorsigera, L. ; Buff. X. lv. (The Cayopollin). A greyish fawn colour ; the circumfer- ence of the eyes and a longitudinal band on the chanfrin brown ; tail marked with black ; size that of the Norway rat. The superior third of the tail furnished with hairs. Did. cinerea, Temm. (The Cinereous Didelphis). A light ash colour, with blackish reflections ; some red on the breast ; the pos- terior half of the tail white ; of the same size as the preceding. From Brazil. Did. murina, L. ; Buff. X. Hi, liii. (The Marmose) J. Fawn- coloured grey ; a brown stripe, in the middle of which is the eye ; tail immaculate : less than a rat. Did. brachyura, Pall., Buff. Supp. VII. lxi. (The Touan). Black, blackish ; flanks of a vivid red ; belly white ; tail shorter than the body. Less than a rat. The three latter species are from South America. Finally, there is one known with palmated feet, which must be aquatic ; it is not ascertained whether or not it has a pouch — it is the * It is the pretended Great Oriental Philander of Seba, of which Linnaeus has made his Did.marsupialus. Buft'on, who has described the male, Supp. III. pi. liii, erro- neously thought the female had no pouch, which was the cause of the improper esta- blishment of a second species, Did. cancrivora. Gm., carcinophaga, Bodd. The Crab- eater is called at Cayenne pian or puant. f Cayopollin, the name of a species that inhabits the mountains of Mexico; it has, somewhat arbitrarily, been applied to this species in particular. X Marmose, a name adopted by Buffon from a typographical error in the French translation of Seba, who assures us in the text that it is called Marmot in Brazil. The truth is, that the Dutch, in the time of Marcgrave, called it Wood-Rat, and the Brazilians Taibi; Rat-de-bois is also its name among the French at Cayenne. Seba must have rendered Bosch-ratte by Marmot. N. B. There has been found, in the plaster quarries near Paris, the fossil skeleton of a Didelphis allied to the Marmose. 110 MAMMALIA. ClIIRONECTES, Itlig*. Did. palmata, Geoff. ; Lutra rnemina, Bodd. ; La petite Loutre de la Guiane, Buff. Supp. III. xxii. Brown above, with three trans- verse grey bands, interrupted in the middle, and white below ; larger than a Norway rat. All the other Marsupialia inhabit eastern countries, New Holland par- ticularly, a land whose animal population seems chiefly to belong to this family. Thylacinus, Temm\. The Thylacini are the largest of this first division. They are distin- guished from the Opossums by the hind feet having no thumb ; a hairy, non-prehensile tail, and two incisors less in each jaw; their molars are of the same number. They consequently have forty-six teeth ; but the ex- ternal edge of the three large ones is projecting and trenchant, almost like the carnivorous tooth of a dog; their ears are hairy, and of a medium size. One species only is known, the Did. cynoeephala, Harris, Linn. Trans. IX. pi. xix, 1, and Ency. Method., Mammif. Supp. pi. vii, f. 3. Size that of a wolf, but stands lower; grey; transverse black stripes on the crupper. It is very carnivorous, and pursues all small quadrupeds. From Van Dieman's Land. Phascogale, Temm. The same number of teeth as the Thylacini, but the middle incisors are longer than the others, and the back molars more bristled, circumstances which approximate them more closely to the Sarigues. They are also allied to them by their small size ; their tail, however, is not prehensile ; their hind thumb, though very short, is still very apparent. Did. penicillata, Shaw, Gen. Zool. I. ii, pi. 113; Schreb. CLII. B. L. Ash-coloured; tail furnished with long black hairs; size that of the Norway rat : lives on the trees in New Holland, and pur- sues insects. Dasytirus minimus, Geoff., Schreb. pi. 152, B. C. (The Dwarf Phascogalis), Scarcely larger than a mouse ; fur soft and reddish. From the south of Van Diemen's Land. Dasyurus, Geoff%. Two incisors and four grinders in each jaw less than the Opossums, so that they have only forty-two teeth; their tail, every where covered with long hairs, is not prehensile. The thumb of the hind foot is reduced to a tubercle, or has even totally disappeared. They are from New Holland, where they feed on insects and dead bodies ; they penetrate into houses, * Chironectes, i. e. swimming with hands. f Thylacinus, from Thulacos, purse. A species of Thylacinus has also been found in the plaster quarries of Paris. X Dasyurus, hairy tail. See Mem.de M.Geoff., Ann. du Mus. III. p. 353, and XV. p. 301. MARSUPIALIA. Ill where their voracity is very inconvenient, &c. Their month is not so wide, their muzzle not so pointed as those of the Opossums ; their hairy ears are also shorter. They do not climb trees. Did. itrsina, Harr. Linn. Trans. IX. xix, f. 2, and Encycl., Supp. f. 6. (The Ursine Opossum). Long rough black hairs, with some irregularly placed white spots; the tail half as long as the body, al- most naked underneath. Inhabits the north of Van Dieman's Land, and is nearly the size of the badger. Das. macrourus, Geoff., Peron. Voy. pi. xxxiii, Schreb. CLII, B, a. (The Long-tailed Dasyurus). Size of a cat ; tail as long as the body ; fur brown, spotted with white, both on the body and tail. The tubercle of the thumb is still well marked in this species, but in the following ones it can no more be seen. Das. Maugei, Geoff., Voy. de Freycin. Zool. pi. iv, Schreb. CLII. B, b. (A kind of olive colour, spotted with white ; no spot on the tail; a little smaller than the preceding. Did. viverrina, Shaw, Gen. Zool. CXI ; White, Bot. Bay, App. 285 ; Schreb. CLII, B, c. Black, spotted with white ; no spots on the tail ; a third less than the first. Perameles, Geoff*. — Thylacis, IUig. The thumb of the hind foot short, like the first Dasyuri, and the two following toes united by the membrane as far as the nails ; the thumb and the little toe of their fore feet are simple tubercles, so that there seem to be but three toes. They have ten incisors above, the external ones sepa- rate and pointed, and only six below; but their molars are the same as in the Opossums, so that they have forty-eight teeth. Their tail is hairy, and not prehensile. The great claws of their fore feet announce their habit of digging in the earth ; and the tolerable length of their hind ones, a swiftness of gait. P. nasutus, G., Ann. du Mus. IV. The muzzle much elongated; ears pointed; fur a greyish brown. At the first glance it resembles a TenrecJ. The species belonging to the second subdivision of the Marsupialia have two broad and long incisors in the lower jaw with pointed and trench- ant edges sloping forwards, and six corresponding ones in the upper jaw. Their superior canini are also long and pointed, but all their inferior ones consist of teeth so small that they are frequently hidden by the gum ; they are sometimes altogether wanting in the lower jaw of the last subgenus. Their regimen is chiefly frugivorous ; consecpiently, their intestines, the caecum particularly, are longer than in the Opossum. The thumb is very large in all of them, and so widely separated from the toes that it seems to slant backwards almost like that of birds. It has no nail, and the two * Pera, purse> Meles, Badger. See Mem. Geoff., Ann. du Mus. torn. IV. t The Peramele Bougainville of Quoy and Gaynard does not differ specifically from the nasutus. The Perarn obesula, Geoff, is not so authentic. ] 12 MAMMALIA. following toes are united by the skin as l'ar as the last phalanx. It is from this circumstance that these animals have received the name of Pha- langers*. Phalangista. Phalangista, Cuv. — Balantia, IUig\. The true Phalangers have not the skin of the flank extended; four back molars in each jaw, with four points in two rows; in front a large one, conical and compressed, and between it and the superior canine are two small and pointed ones, to which correspond the three very small lower ones, of which we have just spoken. Their tail is always prehensile. The tail in some of them is in a great measure scaly. They live on trees in the Moluccas, on which they seek insects and fruit. At the sight of a man they suspend themselves by their tail; and if he gaze at them steadily for some time, he causes them to fall through lassitude. They diffuse a very unpleasant odour, notwithstanding which their flesh is eaten. There are several of them known, of various sizes and colours, all of which are embraced under the Didelphis orientaUs of Linnaeus. M. Temminck thinks he can separate them into species as follows : Ph. ursina, T. (The Ursine Phalanger). Nearly the size of the civet ; fur close, and of a blackish-brown ; the young ones a fawn- coloured brown. From the woods of the island of Macassar. Ph. chrysorrhous, T. (The Golden- cruppered Phalanger). Size of a large cat ; fur of an ash brown ; white beneath ; a golden fawn colour on the croup. From the Moluccas. Ph.maculata, T. ; Buff. XIII. pi. ii; Voy.de Freycin, pi. vii; Voy. du Duperr. pi. iv. (The Spotted Phalanger). Size of a cat; whitish, irregularly spotted or marbled with brown. Ph. cavifrons, T. ; Buff. pi. x, the female ; and Voy. de Duperrey, the male. (The Hollow-fronted Phalanger). The male white ; the female fawn-coloured, with a brown stripe along the back. To these we must add Ph. Quoy, Voy. de Freycin., pi. vi. (The Quoy Phalanger). A greyish-brown ; a blackish-brown longitudinal band on the croup ; top of the head a cinnamon-red; cheeks, throat, and breasts white J. In others, which have hitherto been found in New Holland only, the tail is hairy to the tip. Ph. vulpina ; Did. lemurina and vulpina, Shaw ; Bruno of Vicq. dAz. ; White, Voy. 278. (The Fox-like Phalanger). Size of a stout cat ; greyish-brown, paler beneath ; tail nearly all black. * The name of Phalanger was given by Buffon to two individuals lie had observed, on account of the union of the two toes of the foot. That of Philander is not, as might be thought, derived from the Greek, but from the Malay word Pe/andor, which means Rabbit, applied by the inhabitants of Amboyna to a species of Kanguroo. Seba and Brisson have used it indiscriminately for all the pouched animals. The Phalangers, in the Moluccas, are called Couscous or Coussous. The earlier travellers not having properly distinguished them from the Sarigues, gave cause to believe that this last genus was common to the two continents. f Balantia, from Balantion, purse or pouch. X A very distinct species. % MARSUPIALIA. 113 Ph. CooJcii, Cook's last Voy., pi. viii. (The Phalanger of Cook). Less than a cat; brown above, white underneath; head and flanks red; posterior third of the tail white. Ph. Bougainvillii. (The Phalanger of Bougainville). Size of a squirrel ; ash-coloured above, white underneath ; the posterior half of the tail black; posterior half of the ear white.* Petaurus, Shaw. — Phalangista, Illig. The Flying Phalangers have the skin of the flanks more or less ex- tended between the legs, like the Flying Squirrels among the Rodentia, which enables them to sustain themselves for a few moments in the air, and make greater leaps. They also are only found in New Holland. Some of the species have inferior canines, but they are very small. Their superior canines, and their three first molars, above and below, are very pointed; each of their back molars has four points. -j- Ph.pygmcea; Did. pygmcea, Shaw, Gen. Zool. pi. 114; Schreb. CLXI V, A. (The Flying Dwarf Phalanger). Of the colour and nearly the size of a mouse ; the hairs of the tail regularly arranged on its two sides like the barbs of a quill. Other species have no inferior canini, while the superior ones are very small. Their four back molars present four points, but they are slightly curved into a crescent, which is very nearly the form of those of the Ru- minantia. In the front there are two above and one below, less compli- cated. By this structure they are rendered still more frugiverous than all the preceding species. Ph. petaurus, Shaw, Gen. Zool., pi. cxii; White, Voy. 288. (The Great Flying Phalanger). Resembles the Taguan and the Galeo- pithicus in size ; its fur is soft and close ; its tail long and flattened ; brownish-black above, white beneath. They are of various shades of brown ; some are variegated, and others perfectly white. Ph. sciurea, Shaw, pi. cxiii, 3. (The Bordered Flying Phalan- ger). Size of the brown rat; ash-coloured above, white beneath ; a brown line commencing on the chanfrin and running along the back; edges of the lateral membrane brown; tail tufted, and of the length of the body; its posterior portion black. From the islands near New Guinea. P, peronii, Desm. (The Hairy-footed Flying Phalanger). A reddish-grey ; front of the ears and under part of the body whitish ; toes very hairy and brown ; tail black, longer than the body, and white at the end. Ph. macroura, Shaw, pi. cxiii, f, 2. (The Long-tailed Flying Phalanger). A deep brown above, white beneath ; size of the brown rat ; tail slender, about half as long again as the body. Our third subdivision has the incisors and superior canines and the two * A new species brought to France by M. de Bougainville from his last expedition, f It is of this first division that Desmarcts has made his genus Acrobate. VOL. I. I * 114 MAMMALIA. toes united to the hind feet of the second ; but the posterior thumbs and inferior canini are wanting. It contains but a single genus. Hypsiprymnus, Illig.* The Potoroos are the last animals of this family which retain any trait of the general characters of the Carnaria. Their teeth are nearly the same as thase of the Phalangers, and they still have pointed canines above. The two superior middle incisors are pointed, and longer than the others; the inferior ones are but two, and project forwards. In front they have a long trenchant denticulated molar, followed by four others, bristling with four blunt tubercles. What particularly distinguishes these animals is their hind legs, which are much larger in proportion than the fore ones, that have no thumbs, and the two first toes united as far as the nail ; so that, at a first glance, it seems as though there were but three toes, of which the inner one has two nails. They frequently walk upon two feet, at which times they employ their long and strong tail to support themselves. They have then the form and habits of the Kanguroos, from which they only differ in their superior canine tooth. They are frugivo- rous ; their stomach is large, divided into two sacs, and has several infla- tions ; but their caecum is rounded, and of a middling size. Hyps, minor; Macropus minor, Shaw; White, Bot. Bay, 286; Voy. de Freycin. pi. 10. (The Kanguroo Rat.) Size of a small rabbit; of a mouse-grey. From New Holland, where it is called Po- toroo. It is the only species known. The fourth subdivision only differs from the third in the absence of all canines whatsoever. These are — Macropus, Shaw — Halmaturus, Illig. f The Kanguroos, which present all the characters we have just assigned to the preceding genus, except that the superior canine is wanting, and that their middle incisors do not project beyond the others. The inequa- lity of their legs is still greater, so that on all fours they can only walk slowly and with difficulty; they make vigorous leaps, however, on their hind feet, the great middle nail of which (almost in the shape of a hoof) also serves them for purposes of defence : for, by supporting themselves on one foot and their enormous tail, they can inflict a severe blow with the foot which is at liberty. They are very gentle herbivorous animals, their grinders presenting mere transverse ridges. They have five teeth in all, the front ones being more or less trenchant, and falling out with age ; so that in old kanguroos we frequently find but three. Their sto- mach consists of two long sacs, that are inflated at several places, like a colon. The caecum, also, is large, and has inflations. The radius al- lows a complete rotation of the fore-arm. In these two genera the penis is not bifurcated, but the female organs of generation are similar to those of other Marsupialia. Ipsiprumnos; i. e. raised behind. \ Halmaturus, tail fit for leaping. MARSUPIALIA. 115 M. major, Shaw; Didelphis gigantea, Gm. ; Schreb. CLIII. (The Greater Kanguroo). Sometimes six feet in height. It is the largest of the New Holland animals, was discovered by Cook, in 1779, and is now bred in Europe. Its flesh is said to resemble ve- nison. The young ones, which at birtli are only an inch long, pass into the maternal pouch, even when they are old enough to graze, which they effect by stretching out their necks from their pouch, while the mother herself is feeding. These animals live in troops, conducted by the old males. They make enormous leaps. It ap- pears that we have hitherto confounded, under this name, several species of New Holland and its neighbouring countries, whose fur, more or less grey, only varies by a trifling difference of shade.* There is another species much more anciently known : — M. Brunii; Did. Brunii, Gm. ; Schreb. CLIII. ; called Pelan- dor Aroe, or the Aroe Rabbit, by the Malays of Amboyna. (The Kanguroo of Aroe). Larger than a hare ; brown above, fawn-co- loured beneath. Found in the islands near Banda, and in those of Solor. European naturalists had not paid sufficient attention to the descriptions of the above species given by Valentine and Le Bruyn. M. elegans; Halma. elegans, Per. Voy.. t. xxvii. (The Elegant Kanguroo). Size of a large hare ; transversely striped with brown on a greyish-white ground. Found at the island of St. Peter (a). The fifth subdivision has two long incisors, without canines, in the lower jaw; in the upper, two long incisors in the middle, a few small ones on the sides, and two small canines. It comprehends but one genus. * M. Geoff, distinguishes the Kanguroo enfume, in which the grey is deeper; the Kanguroo a moustaches, which has some white on the front of the upper lip; the Kan- guroo a cou roux, a little less than the others, with some red on the nape of the neck. Messrs. Lesson and Garnot also describe a brown kanguroo, which they call Ouala- bate, Voy. de Freycin. pi. ix. We shall also probably be obliged to make new spe- cies of the Kanguroo roux-cannelle, (K. laniger, Quoy and Gaym.) Voy. de Freycin. pi. ix., and of the Kanguroo cendre-bleuatre ; but all these quadrupeds require to be examined at various ages; and we must ascertain the influence of age and sex upon their colours, previous to a final establishment of the species. g^ (a) This kanguroo is stiled by the traveller mentioned in the text (Peron), Mar opus Fasciatus, the Fasicated Kanguroo. He gives a most interesting account of the intelligence and affections of the females, as displayed towards the offspring, which in early life occupy so peculiar a position externally, to the body of the mother. The young of the opossumus have to undergo a similar process with that of the kan- guroos, and are received, at an early period of gestation, into the external pouch. The time at which the transfer takes place to the pouch is not yet ascertained; but the naturalists, who have carefully examined this subject, have found, that, when the young is seen first attached to the nipple, there is no face, and the nipple seems only to adhere to a round hole in the muzzle of the imperfect offspring. After this, the lips and jaws grow upon the nipple ; until at length, about half an inch of it remains in the young creature's mouth. A keeper of a kanguroo in France, in the employ- ment of an ex-French princess, made notes of the gestation of a kanguroo, from which it appeared that the period had continued from the 6th of May to the Cth of October; and the young kanguroo did not finally quit the pouch till the following January. Several kanguroos have thrived and bred readily in the Zoological Gar- dens, where they are fed on grain, various common vegetables, and hay. — Eng. Ed. i2 I1G MAMMALIA. Koala, Cud. — Lipurus, Gold. — Phascolarctos, Bla'in. The Koalse have a short, stout body, short legs, and no tail. The toes of their fore feet, five in number, when about to seize any object, separate into two groups; the thumb and index on one side, and the remaining three on the other. The thumb is wanting on the hind foot; the two first toes of which are united, like those of the Phalangers and the Kan- guroos. One species only is known : — K. cinerea; Lipurus cinereus, Gold.; Schreb. CLV, A, a. (The Koala). Ash-coloured ; passes one part of its life on trees, and the other in burrows, which it excavates at their foot. The mother car- ries her young one for a long time on her back. Finally, our sixth division of the Marsupialia, or the Phascolomys, Geoff.* Consists of animals which are true Rodentia, as respects the teeth and intestines, their only relation to the Carnaria consisting in the articula- tion of their lower jaw ; and, in a rigorously exact system, it would he ne- cessary to class them with the Rodentia. We should even have placed them there, had we not been led to them by a regular uninterrupted series, from the Opossums to the Phalangers, from the latter to the Kanguroos, and from the Kanguroos to the Phascolomys; and, finally, were it not that the organs of generation are every way exactly similar to those of the Marsupialia. They are sluggish animals, with large, flat heads, short legs, and bodies that look as if they had been crushed, without a tail ; have five nails on each of the fore feet, and four, with a small tubercle in place of a thumb, on each of the hind ones, all very long and fit for digging. Their gait is excessively slow. They have two long incisors in each jaw, almost simi- lar to those of the Rodentia ; and each of their grinders has two trans- verse ridges. They feed on grass ; their stomach is pyriform, and their ca?cum short and wide, furnished, like that of man, and of the ourang-outang, with a vermiform appendage. The penis is bifurcated, like that of the opossums. One species only is known, the Phas. ursinus; Didelphis ursina, Shaw; Peron. Voy. pi. xxxviii, and called by naturalists the Wombat. Size of a Badger; fur abun- dant, of a more or less yellowish brown. It is found in King's Island, to the south of New Holland, where it lives in its burrow, and multi- plies with facility in Europe. Its flesh is said to be excellent. -j- * Phascolomys, a pouched rat, from phaskolon and'mus. f M. Bass has described an animal, externally similar to the Phascolomys, and to which he also gives the name of Wombat, but which has six incisors, two canines, and sixteen molars in each jaw. If there is no erroneous combination of the two dif- ferent descriptions, it will form an additional subgenus, to place near the Perameles. Illiger has already established it under the name of Amblotis, from amblotus, abortus. See Petersb. Mem, 1803—180(5, p. 444, and the Bulletin des Sc. No. 72, An. XI. RODENTJA. 11? ORDER V RODENTIA. We have just seen, in the Phalangers, canini so very small, that we cannot consider them as such. The nutriment of these animals, accord- ingly, is chiefly derived from the vegetable kingdom. Their intestines are long, and their cascum ample ; and the kanguroos, which have no ca- nini whatever, subsist upon vegetables only. The Phascolomys might stand first in that series of animals of which we are about to speak, and which have a system of mastication still less complete. Two large incisors in each jaw, separated from the molars by an empty space, cannot seize a living prey nor tear flesh ; they cannot even cut the food, but they serve to file, and, by continued labour, to reduce it into se- parate molecules — in a word, to gnaw it; hence the term Rodentia, or Gnawers, which is applied to animals of this order. It is thus that they successfully attack the hardest substances, frequently feeding on wood and the bark of trees. The more easily to accomplish this object, the incisors have no thick enamel except in front, so that their posterior edges wear- ing away faster than the anterior, they are always naturally sloped. Their prismatic form causes them to grow from the root as fast as they wear away at the edge; and this tendency to increase in length is so powerful, that if one of them be lost or broken, its antagonist in the other jaw, hav- ing nothing to oppose or comminute, becomes developed to a most mon- strous extent. The lower jaw is articulated by a longitudinal condyle, in such a way as to allow of no horizontal motion except from back to front, and vice versa, as is requisite for the action of gnawing. The molars also have flat crowns, whose enamelled eminences are always transverse, so as to be in opposition to the horizontal motion of the jaw, and to serve the better in trituration. The genera in which these eminences are simple lines, and the crown is very flat, are more exclusively frugiverous ; those in which the emi- nences of the teeth are divided into blunt tubercles are omnivorous ; while the small number of such as have no points more readily attack other ani- mals, and approximate somewhat to the Carnaria. The form of the body in the Rodentia is generally such, that the hinder parts of it exceed those of the front; so that they rather leap than walk. In some of them this disproportion is even as excessive as it is in the Kanguroos. The intestines of the Rodentia are very long; their stomach simple, or 118 MAMMALIA. but little divided ; and their caecum is often very voluminous, even more so than the stomach. In the subgenus Myoxus, however, this intestine is wanting. In the whole of this class the brain is almost smooth and without con- volutions ; the orbits are not separated from the temporal fossae, which have but little depth, and the eyes are altogether directed laterally. The zygomatic arches, thin and curved below, indicate the weakness of the jaws; the fore-arms have nearly lost the power of rotation, and their two bones are often united ; in a word, the inferiority of these animals is visi- ble in most of the details of their organization. Those genera, however, which possess stronger clavicles, have a certain degree of dexterity, and use their fore feet to convey their food to the mouth. Some of them even climb trees with facility ; such are the Sciurus, Lin. The Squirrels, which are distinguished by their strongly compressed inferior incisors, and by their long tail furnished with hairs. They have four toes before and five behind. The thumb of the fore foot is sometimes marked by a tubercle. They have in all four grinders, variously tuber- culated, and a very small additional one above in front, that falls at an early period. The head is large, and the eyes projecting and lively. They are light and active animals, living on trees, and feeding on fruits. Sciurus, Cuv. In the Squirrel, properly so called, the hairs of the tail are arranged on the sides, so as to resemble a large feather. There are a great many species in the two continents. Sc. vulgaris, Buff. VII. xxxii, Schreb., pi. 212. (The Common Squirrel). The back of a lively red ; belly white ; ears terminated by a tuft of hair. Those of the north become of a beautiful ash-blue colour on the back during winter, and at this period supply the fur known by the name of Petit-Gris (miniver), when stripped from the back, and vair when the white part of the belly is attached to the portion from the back. The American species have no pencils to their ears. Such are Sc. cinereus, L. ; Petit-Gris of BufF. X. xxv. (The Grey Squir- rel of Carolina). Larger than that of Europe ; ash-coloured, with a white belly (a). Sc. capistratus, Bosc. Sc. cinereus, Schreb. CCXIII, B. (The Masked Squirrel). Ash-coloured; head black; muzzle, ears, and belly white. Both these species vary in being more or less brown — they are sometimes entirely black. 85T («) There is an error in this description. The Sc. cinereus is the cat sqirrel, which is cinereous above; white beneath; with a tail less distichous than that of other species, longer than the body, and striped with black; inhabits the northern and middle states of America. The animal described by Cuvier as S. cinerius is the S. Carolinensis, the Little Grey Squirrel. — Eng. Ed. RODENTIA. 119 The greater number of the species belonging to the eastern con- tinent are also destitute of these pencils. One of the most beautiful is the Sc. maximus and macrourus, Gm. ;* Buff. Supp. VII. Ixxii. (The Large Squirrel of India). Nearly the size of a cat; above black; the flanks and top of the head of a beautiful bright maronne ; the head, and all the under parts of the body, and the inside of the limbs pale yellow ; a maronne-coloured band behind the cheek. It inhabits the palms, and is extravagantly fond of the milk of the cocoa-nut. There are, also, several Squirrels in warm climates, that are re- markable for the longitudinal bands which vary their fur. Such are the Sc. getulus, L. ; Buff. X. xxvi. (The Barbaresque). The bands of which extend to the tail, and even on it. Sc. palmarum, L. : Buff. X. xxvi. (The Palmist). On which the stripes are confined to the back. It is probable that we shall have to separate from the squirrels certain species which have cheek-pouches, like the Hamsters, and pass their lives in subterraneous holes, the Tamice of Illiger. For instance the Sc. striatus, L. ; Buff. X. xxviii. (The Ground Squirrel). Which is found throughout all the north of Asia and America, particularly in the pine forests. The tail is more scantily supplied with hairs than that of the European Squirrel; the ears smooth, and skin brown, with five black stripes and two white ones. We ought, also, most probably, to distinguish the GuerUnguets, a spe- cies with a long and almost round tail, and an enormous pendant scrotum. They are found in both continents-}-. The following have been separated already. Pteromys, Cuv\. Or the Flying Squirrels, to which the skin of the flank, extending be- tween the fore and hind legs, imparts the faculty of supporting themselves for some moments in the air, and of making very great leaps. There are long bony appendages to their feet, which support a part of this lateral membrane. There is a species in Poland, Russia, and Siberia. Sc. volans, L. ; Schreb. CCXXI1I. (The Flying Squirrel). Ash- coloured, grey above; white underneath; size of a rat; the tail only half the length of the body. It lives solitarily in the forests. One from North America. Sc. voluccella, L. ; Buff. X. xxi. (The American Flying Squir- rel). Reddish-grey above ; white beneath : size less than that of * A comparison of the figure of Pennant with that of Sonnerat is sufficient to prove that they represent the same animal. f We have found, however, in the Tamice and GuerUnguets, the same kind of teeth as in the Squirrels and Pteromys. * Pteromys, Winged Rat. 120 MAMMALIA. the preceding; tail three-fourths as long as the body. It lives in troops in the prairies of the temperate regions of North America. There is one found in the Indian Archipelago, that is nearly the size of a cat; the male is of a fine lively maronne above, and red underneath ; the female brown above, and whitish underneath. It is- the "ioc. petaurista, L. ; Buff. Supp. III. xxi, and VII. lxvii. (The TagiKin). The same Archipelago produces another small one, the Sc. sagitta, L. A deep brown above; white beneath; distin- guished from other species, the small ones especially, by its mem- brane, which, as in the Taguan, forms an extremely acute projecting angle behind the tarsus. •M. Geoffroy has very properly separated from this genus the Cheiromys, Cuv. % Or the Aye-Ayes, whose inferior incisors, much more compressed, and, in an especial manner, more extended from front to back, resemble plough- shares. Each foot has five toes, of which four of the anterior are exces- sively elongated, the medius being more slender than the others ; in the hind feet the thumb is opposable to the other toes ; so that they are in this respect among the Rodentia, what the Opossums are among the Car- naria. The structure of their head is otherwise very different from that of the other Rodentia, and is related to the Quodrumana in more points than one. There is only one species of the Aye-Aye known. It was disco- vered at Madagascar by Somierat. It is the Cheir. Madagascarien- sis; Sc. Madagascar., Gm. ; Buff. Supp. VII. lxviii. (The Aye- Aye). Size of a hare, of a brown colour, mixed with yellow; tail long and thick, with stout black bristles ; ears large and naked. It is a nocturnal animal, to which motion seems painful; it burrows under ground, and uses its slender toe to convey food to its mouth. Linnseas and Pallas united in one single group, under the name of Mus. Lin. All the Rodentia furnished with clavicles, which they could not distin- guish by some very sensible external character, such as the tail of the squirrel or that of the beaver, from which resulted the utter impossibility of assigning to them any common character; the greater number had merely pointed lower incisors, but even this was subject to exceptions. Gmelin has already separated from them the marmots, dormice, and the jerboas; but we carry their subdivision much further, from considerations founded on the form of their grinders. Arctomys,-j~ Gm. The Marmots, it is true, have the inferior incisors pointed like those of the greater number of animals comprehended in the great genus Mus; * Cheiromys, a rat with hands. f Arctomys, Bear Rat. RODENTIA. 121 but their grinders, like those of the squirrel, amount to five on each side above, and four below, all bristled with points ; accordingly, some species are inclined to eat flesh, and feed upon insects, as well as grass. There are four toes and a tubercle in place of a thumb to the fore feet, and five toes to the hind ones. In other respects these animals are nearly the direct reverse of the squirrels, being heavy, having short legs, a middle- sized or short hairy tail, and a large flat head, passing the winter in a state of torpor, and shut up in deep holes, the entrance of which they close with a heap of grass. They live in societies, and are easily tamed. Two species are known in the eastern continent. Arct. alpinus; Mus. alpinus, L. ; Buff. VIII. xxviii. (The Al- pine Marmot). Large as a hare ; tail short ; fur yellowish-grey, with ash-coloured tints about the head. It lives in high mountains, immediately below the region of perpetual snow. Arc. bobac; M. bobac, L. ; Pall. Glir. V; Schreb. CCIX. (The Bobac). Size of the preceding; of a yellowish-grey, tinted with red about the head. Inhabits low mountains and hills, from Poland to Kamschatka, and frequently digs its burrow in the hardest soil.* America also produces some species. Arct. monax, Buff. Supp. III. xxviii. (The Maryland Marmot). Grey ; tail blackish, as well as the top of the head. Arct. empetra, Pall.; Schreb. CX. Less than the preceding; grey ; red beneath. Spermophilus, Fred. Cuv. We apply this name to those Marmots that have cheek pouches. The superior lightness of their structure has caused them to be called Ground Squirrels. Eastern Europe produces one species. M. citillus, L. ; Buff. Supp. III. xxxi. (The Souslik or Zizel). A pretty little animal, of a greyish brown, watered or mottled with white, the spots very small, which is found from Bohemia to Siberia. It has a peculiar fondness for flesh, and does not spare even its own species. North America has several species of them, one of which is re- markable by the thirteen fawn-coloured stripes which extend along the back on a blackish ground. It is the Thirteen-striped Souslik, Arct. 13-Mneatus, Harl. ; or Sciurus lS-lineatus, Mitchell; or Arct. Hoodii, Sabine, Lin. Trans. XIII. pi. xxixj~. There is one of the Rodentia which it appears we must approximate to the Marmots, that is remarkable for living in large troops in immense burrows, which have even been styled villages. It is called the Prairie Dog, or Barking Squirrel, the latter appellation arising from its voice, * Russian travellers in Bucharia mention some other Marmots, Arct. fulvus, Arct. Upto-dactylus, Arct. musogaricus, which are not yet perhaps sufficiently distinguished from the Boubak or from the Souslik. f Add Arct. Parrii, Richards. App. Parry's Voy. — Several of the Marmots an- nounced in the travels of Lewis and Clarke, Parry, Franklin, &c. Arct. Franklinii, Richardsonii, pruinosa, seem to belong to this subgenus. See Sabine, Lin. Trans. XIII. pi. xxvii, xxviii, &c. 122 MAMMALIA. which resembles the bark of a small dog. It is the Arct. ludovicianus of Say, Jour, to the Rocky Mountains, I. 451. M. Rafinesque, who states it has five toes to each foot, makes it the type of his genus Cynomys. Myoxus, Gm.* The Dormice have pointed lower incisors, and four grinders, the crown of each of which is divided by enamelled lines. They are pretty little animals, with soft fur, a hairy and even tufted tail, and a lively eye, which live on trees like squirrels, and feed on fruit. Of the numerous order of the Rodentia, this is the only subgenus in which there is no caecum. They become torpid in winter like the Marmots, and pass through it in the most profound lethargy j. M.glis,~L.; Buff. VIII. xxiv. (The Fat Dormouse) (a). Size of of a rat ; ashy grey-brown above, whitish underneath ; of a deeper brown around the eyes; tail very hairy the whole of its length, and disposed somewhat like that of a squirrel, and frequently a little forked at the extremity. It inhabits the south of Europe, and nestles in the hollows of trees and fissures of rocks. It sometimes attacks small birds. This is probably the rat fattened by the ancients, among whom it was considered a delicacy of the very highest de- scription J. M. nitela, Gm. ; Buff. VIII. xxv. (The Garden Dormouse). Somewhat less than the preceding; greyish-brown above; white un- derneath; black round the eye, which extends to the shoulder; tail tufted and black, tuft white. Common in the gardens of Europe, where it shelters itself in holes about the walls, and does much in- jury to trees. M . avellanarius, L. ; Buff. VIII. xxvi. (The Common Dor- mouse). Size of a mouse ; cinnamon-red above ; white beneath ; hairs of the tail somewhat disposed like a feather. From the forests of all Europe. It constructs its nest of grass on low branches, to bring up its young; the rest of the time, and particularly during winter, it remains in the hollows of trees. § * Myoxus, Rat with a pointed nose. f So natural is this to them, that a dormouse from Senegal (M. Coupeii), which had never experienced it in its native country, fell into a profound sleep in Europe the moment it was exposed to the cold. X The M.dryas of some authors (Schreb. 220, B) does not appear to me to differ from the Fat Dormouse. § Add Myoxus Coupeii, Fred. Cuv. Mammif. ggp° (a) The instinct of the dormouse in providing for itself a proper retreat dur- ing the period of hybernation (see note, p. 71 of this volume) is often displayed with a prescience and circumspection which are scarcely credible in an animal usually rated at the lowest possible estimate in the scale of intelligence. A French natu- ralist has placed on record, in the Bibliotheque Universelle, an anecdote relating to a dormouse, which is at once curious and instructive. He placed four of these ani- mals in a cold temperature, which soon brought them into a state of lethargy, with the exception of one, which escaped secretly from the apartment. Some time after- wards, it was found in a deep cellar in the same house, where it had dug up the earth, and scraped the neighbouring wall, in order to heap up the mould and plaster, so as to form a mound of two feet in size. This mound was raised near a situation where RODENTIA. 123 We should place near the Dormice, the Echimys, Geoff. — Loncheres, Illig . Four grinders also, but formed in a peculiar way ; the upper ones con- sisting of two blades, bent into the shape of a V, and the under ones of one blade only that is bent, and of another that is simple. The fur of several species is very rough, and intermixed with flattened spines or prickles, like sword blades. From America. One of them, Ech. chrysuros; Hystrix chrysuros, Schreb. CLXX, B; Lerot a queue doree, Buff. Supp. VII. 72. (The Golden-tailed Echimys). More than twice the size of the brown rat ; it is a beautiful animal, of a chestnut-brown colour ; white belly ; an elongated crest of hairs, and a white longitudinal band on the head ; the tail is long and black ; its posterior half is yellow. From Guiana. Ech. rufus; Rat epineux, Azzara, Voy. pi. xiii. (The Red Echi- mys). Size of a rat, and of a reddish grey ; tail shorter than the body. It is found in Guiana, Brazil, and Paraguay. It excavates long subterraneous galleries. Others, again, have merely the ordinary kind of hair, more or less rough. The most remarkable is the Ech. dactylicus, Geoff. (The Long-toed Echimys). Which is still larger than the golden-tailed one, and has the two middle toes of the fore feet double the length of the lateral ones. Its scaly tail is longer than the body ; its fur is a yellowish grey, and the hairs on its nose form a crest directed in front.* Hyrdomys, Geoff. The Hydromys have many external points of relation to the Echimys, but they are distinguished from all other rats by their hind feet, two thirds of which are palmated ; their twe molars have also a peculiar character in the crown, which is divided into obliquely quadrangular lobes, whose sum- mits are hollowed out like the bowl of a spoon. They are aquatic. * Add the Echimys of Cayenne, the Silky Echimys. I suspect the Mus. para- doxus, Thomas, Lin. Trans. XI. (Heteromys, Lesson,) differs from the Echimys in its cheek-pouches only. However, not having seen its teeth, I cannot arrange it. a door admitted, between its lower edge and the floor, a current of air; and, in order to get rid of the inconvenience, the dormouse had previously fixed up a piece of board, which it absolutely detached from a shelf, and placed against the door. But this was not all. The dormouse, it was found, had untied a straw rope which encir- cled some bottles that lay in the cellar; of this it made a bed, which it lastly sur- rounded with a rampart curiously and ingeniously constructed; for this wall of se- curity was composed of the fragments of the bottles literally broken for the purpose of being placed as a wall of separation between the bed of the dormouse and the rats that might chance to invade it. The dormice are found in great numbers, in bur- rows, on the highest of the rocks of the Alpine mountains. They come out in wet weather, and generally announce the approach of rain by a shrill, and very peculiar whistle. The inhabitants of the Alp's regard their appearance abroad as faithful indications of the weather. Dr. M. Hall, in the paper on hybernation already alluded to, states, that dormice, supplied with cotton wool, make themselves nests, and be- come lethargic. — Eng. Ed. 124 MAMMALIA. Several of these animals have been sent to Europe from Van Dieman's land, some of which have the belly white, and others fawn- coloured, but all of a deep brown above, with a long tail, which is black at the base, and the posterior half white. They are sometimes double the size of the brown rat. Hydromys leucogaster, and Hyd. chrysogaster, Geoff. An. Mus. VI. pi. xxxvi. Capkomys, Desmar. The Houtias have four molars every where with flat crowns, the enamel of which is folded inwards so that it forms three angles on the external edge, and a single one on the internal edge of the upper teeth, and the inverse in the lower ones. The tail is round and scantily philose ; they have, like the rats, five toes to the hind foot, and four, with the rudiment of a thumb, to the fore feet ; their form is that of a rat ; as large as a rabbit or hare. Two species are known. Cap. fouvnieri Desmar., Mem. de la Soc. d'Hist. Nat. de Par. I. 1823. (The Congo Houtia)*. Muzzle brown; the under part of the neck whitish ; tail brown, but half the length of the body. Cap. prehensilis, Pcessig. ; Houtia Carvalli. Less than the pre- ceding ; brown, with a whitish throat ; tail red, as long as the body, and partly naked at the end. Both species inhabit the island of Cuba, and, together with the Agoutis, at the time of the discovery, constituted the principal game of the inhabitants. Mus. Cuv. The Rats, properly so called, have three molars, of which the anterior is the largest; its crown is divided into blunt tubercles, which, by being worn, give it the shape of a disk, sloped in various directions ; the tail is long and scaly. These animals are very injurious, from their fecundity, and the voracity with which they gnaw and devour substances of whatever kind. There are three species which have become quite common in our houses, viz. M. musculus, L. ; Buff. VII, xxxix. (The Common Mouse). Known in all times and at all places. M. rattus, L. ; Buff. VII, xxxvi. (The Black Rat). Of which no mention is made by the ancients, and which appears to have en- tered Europe in the middle ages. It is more than double the size of the mouse, in all its dimensions. The fur is blackish. Several in- dividuals have been occasionally found connected by the interlacing of their tails; constituting what the Germans style the King of Rats.-f M. decunumus, Pall. ; Buff. VIII, xxvii. (The Surmulot, or Common Norway or Brown Rat). Which did not pass into Europe till the eighteenth century, and is now more common in Paris and other large cities than the Black Rat itself. It is larger than the * This is the Isodon piloritics, Say. Zool. Journ. No. 2, p. 229. t See Bellerman on the King of the Rats (in German), Berlin, 1820. RODENTFA. 125 latter by one-fourth, and differs from it also by its reddish-brown hair.* These two large species appear to have originated in the East, and have been transported in ships, together with the mouse, to all parts of the globe. Eastern Tartary and China have a Rat equal to the Surmulot. M. caraco, Pallas, Glir. XXIII ; Schreb. clxxvii. (The Caraco Rat). Of a light colour; tail a little shorter than the preceding, and the jaws stronger. There is another in India, one-fourth larger than the Brown Rat, the Rat perehal, Buff. Supp. VII. lxix, which is of a reddish brown. There is a large one in the Indian Archipelago, the M. setifer, Horsf. Jav. Of a blackish brown. These last two species are set with rough bristles, which extend beyond the hairs. One of the largest and most mischievous rats known is the M. pilorides, Pall, and Gm. (The Musk-Rat of the Antilles). Fifteen inches in length, without the tail, which is still longer than the body; hair coarse, of a deep black above, and whitish beneath. 'j~ Fewer species have been observed of the size of the mouse. M. cahirinus, Geoff., Descr. de l'Eg. Mammif. (The Cairo Mouse). Spines on the back, in place of hairs; it is noticed by Aristotle. There is scarcely known in France more than one species which lives remote from houses — the M. sylvaticus. (The long-tailed Field-Mouse). The Mulot of Buff. VII, xli, which is hardly larger than the mouse, and is distinguished from it by its red fur. It does much injury to trees, and sometimes penetrates into gardens. It seems, however, that in some of the provinces there is a smaller and grey species, which has also been observed in England, (M. messorius, Shaw, Vol. II. Part 1, Frontisp.), and a third still more diminutive — the Dwarf Field-Mouse, (M. pumilus, Fr. Cuv. Mam- mif). It remains for me to observe, that there are still numerous discoveries to be made in our country respecting the species of these very diminutive quadrupeds. J * It appears to belong to Persia, where it lives in burrows. It was not till 1727, that, after an earthquake, it arrived at Astracan, by swimming across the Volga. f Pallas and Gmelin erroneously describe it as being entirely white. The earlier historians of the colonies attribute to it the above colours, which are precisely such as we have seen on the animal. X To this division, most probably, belong the M. agrarius, M. minutus, M. sorici- nus, M. vagus, M. betulinus, M. striatus, M. barbarus, Schreb. Here, also, should come the enormous Mus giganteus, Hardw. Linn. Trans. VII. xxviii. There should be likewise added the Radiated Rat of the Cape, M.pumilio, Sparm. the M. cyanus, the Grey-Blue Rat of North America, Molina, and several other spe- cies, some of which are not even mentioned by authors, and others which are de- scribed with too little reference to other species. This is the reason why most of the rats of Azzara cannot be properly classed until they are re-examined. The same ob- servation applies to a great many of the Rodentia of M. Rafinesque. Their descrip- tions are too short to be of any use. \ C 2Q MAMMALIA. Warm climates produce rats similar in every particular to those of which we have just spoken, except that their tails are more hairy* (a). Gerbillus, Desm. — Meriones, Elig. The Gerbils have molars that differ very little from those of rats, merely becoming sooner worn, so as to form transverse elevations. Their superior incisors are furrowed with a groove ; their hind feet are some- what longer in proportion than those of rats in general, and their thumb and little toe slightly separated. Their tail is long and hairy. The sandy and warm paits of the eastern continent produce several species. G. indicus; Dipus indicus, Hardw., Linn. Trans. VIII, pi. vii; Herine, Fred. Cuv. Mammif. (The India Gerbil). Size of the fat Dormouse; fawn coloured above, whitish beneath; tail longer than * Hypudceus variegatus, Lichtenst. var. flava. — Meriones syenenses, Id. ; to which must be added the Arvicola messor, Le Comte, Arv. hortensis, Hark, or Sygmodon, Say; distinguished, however, by hairy ears, like the Otomys. Another group, with hairy tails also, but whose teeth wear away faster, will include the Hypudmis obesus, Lichtenst, the Mus ruficauditus, Id. His Meriories sericeus should form a third, characterized by the projecting ridges of the molars, which al- ternately catch in each other. We then have to group the Neotoma floridanum of Say, or the Arvicola floridanus of Harlan, and the Arvicola gossypina of Le Comte, two rats which, size excepted, are very similar even in their colours, whose teeth, provided with roots, if worn a little, have crowns formed like those of the Arvicola. These animals, however, previous to a definite classification, require to be com- pletely examined and compared, internally as well as without. gggT (a) Amongst the specimens of this genus in the Zoological Gardens in Lon- don, are the Mus Rattus, a species so long known in this country as to be considered aboriginal to it. This rat was formerly very abundant, but for many years has been almost wholly displaced by the Surmulot, M. decumanus. Very lately, however, the M. rat/us has very unaccountably re-appeared in several places in London. A spe- cies not described by Cuvier is to be seen in the gardens in Regent's Park; it is the Gigantic Rat, M. giganteus, and was sent from Bombay by that great patron of sci- ence, the Earl of Clare. Here, also, are to be seen some Barbary mice, the M. bar- barns of Linnaeus; since whose time the species entirely eluded all observation, until it was recently recovered by the Zoological Society. It is a species common in Bar- bary. The Long-tailed Field-Mouse, M. sylvaticus, will also be found in the above collection. The singular nest of the Harvest-Mouse, M. messorius, has been ren- dered too interesting an object of contemplation, by White of Selborne, to be passed over. He found it suspended upon the head of a thistle, in a wheat-field. The nest was of the size of a cricket-ball, was perfectly round, and was composed of blades of wheat, which were platted together with wonderful art. What was most curious was the absence of any aperture in the ball, for exit or entrance; and yet, the particular nest which Mr. White examined contained eight young, which so completely occu- pied the cavity, that it was apparently impossible for the creatures to turn themselves, in order to seize the mother's teat; and still more was it difficult for the mother to find room in the nest. These nests, observes the author of the " British Naturalist," vary in shape, being round, oval, or pear-shaped, with a long neck, and are to be distinguished from those of any other mouse, by being generally suspended on some growing vegetable — a thistle, a bean-stalk, or some adjoining stems of wheat, with which it rocks and waves in the wind; but, to prevent the young from being dis- lodged by any violent agitation of the plant, the parent closes up the entrance so uniformly with the whole fabric, that the real opening is with difficulty found. — Eng. Ed. RODENTIA. 127 the body, and blackish towards the end. To this species should be approximated the G. meridianus ; D, meridianus, Schreb. 231. (The Torrid or Sand Gerbil). Which is about the same colour, but a little smaller. G. tamaricinus; D. tamaricinus, Schreb. 232. (The Tamarisk Gerbil). The tail is annulated with brown. G. pyramidum; D. pyramidum, Oliv. (The Gerbil of the Pyra- mids). The hind feet more elevated; size of the Garden Dormouse; its fur is red above, whitish beneath. There is one in Senegal of a livelier red and a purer white. Another at the Cape, a little larger, reddish, and the tail less hairy at the end. A third in Nubia, nearly half the size, of a light red above, and a beautiful white beneath. The Meriones, Fred. Cuv. The Meriones, which we separate from the other Gerbils, have the hind feet still longer, the tail nearly naked, and a very small tooth fronts the superior molars ; characters which approximate them to the Jerboas. Their upper incisors are grooved, like those of the Gerbils, and their toes also are similar. There is a small species in North America, the Mus canadensis, Penn. ; Dipus canadensis, Sh. II, Part 1, pi. 161; Dipus americanus, Barton. Size of a mouse; fawn-coloured grey ; tail longer than the body. An animal of the greatest agility, that shuts itself up in the earth, and passes the winter in a state of lethargy.* Cricetus, Cuv. The Hamsters have nearly the same kind of teeth as rats, but their tail is short and hairy, and the two sides of their mouth are hollowed, as in certain of the species of Simiae, into sacs or cheek pouches (a), in which they transport the grain they collect to their subterraneous abodes. C. vulgaris; M. cricetus, L.; Marmotte d'Allemagne, Sj-c; Buff. XIII, xiv. (The Common Hamster, or German Dormouse). It is larger than the rat; of a reddish-grey above, black at the flanks and underneath, with three whitish spots on each side. The four feet are white, as are also a spot under the throat, and another under the breast; some individuals are all black. This animal, so pleasingly diversified in its colours, is yet one of the most destructive that lives, on account of the quantity of corn which he collects, and with which he fills his burrow — a receptacle sometimes no less than seven feet deep. It is common in all the sandy regions that extend from the north of Germany to Siberia. * Add Gerbillus labradorius, Harl., or M. labrad., Sabine, Frankl. Voy. p. 661. ggp" (a) The cheek-pouches are bags situated between the cheeks and the jaws of several genera of animals belonging to the Quadrumana, as well as that now un- der consideration. These pouches, which are particularly formed in the Hamsters, are destined to be receptacles for the food which the animals take in, and ultimately chew at their leisure. — Eng. Ed. 128 MAMMALIA. This last country produces several small species of Hamsters, de- scribed by Pallas.* Arvicola, Lacep. The Campagnoles, like the rats, have always three grinders, but with- out roots, each one being formed of triangular prisms, placed alternately on two lines. They may be subdivided into several groups, viz. — Fiber, Cuv. The Ondatras or Field Rats, with semi-palmated hind feet, a long, scaly, and compressed tail, of which one species only is well known. F. vulgaris; Castor zibeticus, L.; Mus zibeticus, Gm. ; Buff. X, i. (The Canadian Musk-Rat or Ondatra). As large as a rabbit, of a reddish-grey. In winter they construct, on the ice, a hut of earth, in which several of them reside together, passing through a hole in the bottom, for the roots of the acorus (a), on which they feed. When the frost shuts up this hole, the musk rats are under the necessity of eating one another. It is this habit of building which has induced some authors to refer the Ondatra to the genus Castor. The second subdivision is that of Arvicola, Cuv. — Hypud.eus, Illig. Our common Field Rats, or ordinary Campagnoles, which have a hairy tail, about the length of the body, and without palmated feet. A. vulgaris; Mas amphibius, L. ; Buff, VII, xliii. (The Water Rat). A little larger than the common rat, of a deep greyish- brown ; tail the length of the body. Inhabits the banks of rivers, and digs in marshy places in pursuit of roots ; but it swims and dives badly (b). A. terrestris; Mus ierrestris, Lin. (The Schermaus, or Digger Rat of the Americans). Seems to differ from the Water Rat only in being somewhat smaller; its tail, also, is shorter. It lives under ground, like the mole, but especially in the meadows of high grounds. It excavates galleries, and transports the earth which it raises from its hole to some distance from the opening. Its magazines, which are principally filled with the roots of the wild carrot, cut into two- inch pieces, are frequently two feet in diameter. * M. accedula. — M. arenarius. — M. phceus. — M. songarus. — M. furunculus. See Pall. Glir. and Sch. @3T ( a ) This is the plant called Sweet Flag, and was considered by Linnaeus to be the only aromatic one which grew in the northern regions. Notwithstanding the certainty of Cuvier's account, yet, it is laid down in Loudon's elaborate work entitled the " Encyclopaedia of Plants," that no cattle whatever eat this plant. — Eng. Ed. $3f° (6) It would appear that the species called the Short-tailed Water-Rat is grega- rious, and that troops of them habitually migrate to places where suitable food maybe obtained. They have been known to be particularly partial to spots where the Eqiii- setum limosum (Smooth Horse-Tail) is abundant. How such large companies as have been frequently observed, can change their localities without being seen in their transits, is a mystery which still remains unsolved. — Eng. Ed. RODENTIA. ]'29 A. arvalis; Mus arvalis, L. ; Buff. VII, xlvii. (The Campag- nol, or Little Field-Mouse, called also Mulot in some provinces, but improperly). Size of a mouse ; of a reddish-ash colour ; tail not so long as the body. It inhabits holes which it excavates in the earth, where it collects corn for the winter. The multiplication of this animal is sometimes so excessive as to cause much injury. A. ceconomicus, Mus ceconomicus, Pall. Glir. XIV, A. ; Schreb. cxc. (The Meadow Campagnol). A little darker coloured, and the tail somewhat shorter. It lives in a sort of oven-shaped cham- ber, dug under the turf, from which several narrow and ramifying ca- nals run in various directions ; other canals communicate with a se- cond cavity, where it accumulates its provisions. From all Siberia. It is thought to have been found in Switzerland and in the south of France, particularly, as we are assured, in the potato fields.* Georychus, niig. Or the Lemmings, Cuv., have very short ears and tail, and the toes of the fore feet peculiarly well formed for digging. The two first species .have five very distinct nails to each of the fore feet, like the rat-moles and the jumping-hares. G. lemmas; Mus lemmas, L. ; Pall. Glir. XII, A. B. Schreb. cxcv. (The Lemming). A northern species, as large as a rat, with a fur varied with black and yellow ; very celebrated in consequence of the migrations which it makes from time to time, at periods alto- gether unsettled, and in bodies infinite in their number. It is said, that, on such occasions, they proceed in a straight line, without any river, mountain, or other obstacle impeding them, and that they de- stroy every thing on their route. Their usual residence appears to be the shores of the Arctic Ocean. G. vulgaris; Mus aspalax, Gm., Pall. Glir. X, Schreb. CCV. (The Zocor). Reddish-grey; the three middle nails of the fore feet long, arcuate, compressed, and trenchant, for cutting earth and roots. The limbs are short; there is scarcely any tail; and the eyes are excessively small. From Siberia; where it always lives under ground, like the mole and rat-mole. It feeds principally on' the bulbs of different liliacea? (plants of the lily tribe). The third species, like the other animals comprised in the great genus of rats, has merely the rudiment of a thumb on the fore feet. G. hudsonius; Mus hudsonius, Gm., Schreb. CXC VI. (The Lemming of Hudson's Bay). A light pearly-ash colour ; without tail or external ears ; the two middle toes of the fore foot of the male seem to have double nails, because the skin of the end of the toe is * Here, most probably, would come the M. saxatilis, alliums, rutilus, 'gregalis, and socialis, Pall. Glir. But the M. lagurus and torquatus come nearer to the Lemmings. There are several field-rats, or campagnols, in North America, such as the Arvicola xanthognatha, Leach, Miscel. I, pi. xxvi. — Arvicola pennsylvanica, Wilson, Amer. Ornith. VI, pi. 1, F. 3. — Arv, palustris, Harl. &c, Better figures, and new and com- parative descriptions of the preceding species, are much wanted. Vol. i. k 1 30 MAMMALIA. callous, and forms a projection under the point of the nail — a struc- ture which has not been met with, except in this animal. It is the size of a rat, and lives under ground, in North America. Otomys, Fred. Cuv. The Otomys are nearly allied to the field rats, and have also three grinders, but they are composed of slightly arcuated laminae, arranged in file.* Their incisors are grooved with a longitudinal furrow, and the tail is hairy, as well as the ears, which are very large. The species known is 0. capensis, Fred. Cuv. (The Cape Oto- mys). Inhabits Africa. Size of a rat; fur annulated with black and fawn colours ; tail a third shorter than the body.-j v Dipus, Gm. The Jerboas \ have nearly the same kind of teeth as. the true rats, ex- cept that there is sometimes a very small one immediately before the up- per molars. The tail is long and tufted at the Cnd; the head large; the eyes large and prominent; but their principal character consists in their posterior extremities, which, in comparison with the anterior, are of a most immoderate length, and above all, in the metatarsus of the three middle toes, which is formed of one single bone, resembling what is called the tarsus in birds. It is from this disproportion of the limbs that they were named by the antients Biped Rats; and in fact they seldom move otherwise than by great leaps on their hind feet. There are five toes to each of the fore feet; and, in certain species, besides the three great toes to the hind feet, there are small lateral ones. They live in burrows, and fall into a deep lethargy during the winter. D.sagitta; M. sagitta, L. ; Buff. Supp. VI. xxxix and xl. The Jerbao has only three toes, and is the size of a rat ; a light fawn co- lour above ; white beneath ; tuft of the tail black, the tip white. Is found from Barbary to the north of the Caspian sea. D. hirtipes, Lichtenstein. (The Hairy-footed Jerboa). The head more compressed than in the others ; only three toes to the hind feet, as in the Jerboa, but they are more hairy. From Africa.§ D. jaculus; M. jaculus, Pall. Glir. XX. Schreb. CCXXVIII. (The Alactaga). Two small lateral toes; ears longer than those of the Jerboa, butjias nearly the same colours. Pallas has observed them of three sizes, from that of a rabbit to that of a rat : they are * They are exact models, in miniature, of the grinders of the elephant. f It is the same animal described and represented in the essay on the genus of rats, by M. Brantz, Berlin, 1827, under the name of Euryotis irrorata. % There has lately appeared an excellent paper on the Jerboas, by M. Lichten- stein, in which that learned naturalist describes and figures ten species. I can only refer my readers to the paper itself. It is inserted in the Journal of the Acad, of Berlin. § Add the Dlp.telurn, D.platurus, and D. lagopus of Eversman, Voy. de Mayen- dorf en Boucarie, p. 390. RODENTIA. 13] probably as many species.* One or the other is found from Bar- bary to the Eastern Ocean, and as far as the north of India. Helamys, F. Cue. — Pedetes, llllg.-f We separate from the other Jerboas, and the whole of the genus Rats, the Jumping Hares, which, like the Jerboas, have a large head, and great eyes, a long tail, and the anterior part of the body extremely small in comparison to the posterior, although the disproportion is much less than in the true Jerboas. The peculiar characters of the Helamys are four grinders everywhere, each one composed of two lamina? ; five toes to the fore feet, armed with long and pointed nails, and four to their great hind ones, all separate, even to the bones of the metatarsus, and terminated by large nails, almost resembling hoofs. This number of toes is the inverse of that most common among the rats. Their inferior incisors are trun- cated, and not pointed like those of the true Jerboas, and of the greater part of the animals comprised under the genus of rats. One species only is known, the H. caffer.; Mus.caffer., Pall. ; Dipuscaffer., Gm., Buff. Supp.VI. xli, and better, Fred. Cuv. Mammif. It is the size of a hare, of a light fawn colour, and has a long tufted tail, with a black tip. In- habits deep burrows at the Cape of Good Hope. S pal ax, Guldenstedt. The Rat-Moles have also been very properly separated from the Rats, although their grinders are three in number, and tuberculous, as in the true rats, and the hamsters, and are merely a little less unequal. Their incisors, however, are too large to be covered by the lips, and the extre- mity of the lower ones is in a sharp edge, and not pointed. Their legs are very short; each foot has five short toes, and as many flat and slender nails. Their tail is very short, or rather there is none; the same ob- servation applies to their external ear. They live under ground like the moles, raising up the earth like them, although provided with much in- ferior means for dividing it ; but they subsist on roots only. S. typhus; M. typhus, Pall. Glir. pi. viii, Schreb. 206. (The Zemni Slepez or Blind Rat-Mole). A singular animal, whose as- pect is utterly misshapen by its bulky head, which is angular on its sides, by its short feet, by the entire absence of a tail ; but, above all, by its possessing no eye which can be seen externally, it having merely under the skin a small black point, which would seem or- ganized for an eye, without being able however to minister to vision, inasmuch as the skin passes over it without either opening or be- coming thinner, and not having in this spot less hair than any other part. It is rather larger than our rat; its fur is smooth, and of an ash colour, bordering on a red. This is the animal, in the opinion of Olivier, to which the ancients alluded when they spoke of the mole as being perfectly blind. * Pallas has latterly distinguished the small Alactagns by the name of Dip. acontion. f Pedeles, Juniper, Helamys, Jumping-Rat. k2 132 MAMMALIA. The islands in the straits of Sunda produce a rat-mole as large as a rabbit, of a deep grey colour, with a white longitudinal stripe on the head, the Spalax javanus. From the rat-moles themselves should have been separated the Bathyergus, lllig. — Orycteres, Fr. Cuv. The Orycteres, which, with the general form, feet, and truncated inci- sors of that genus, have four grinders throughout. Their eye, though small, is visible, and they have a short tail. B.maritimns; Mus maritimus, Gm. ; Taupe des dames, Buff. Supp. VI. xxxviii. (The Maritime Rat-Mole). Nearly the size of a rabbit; the superior incisors furrowed with a groove, and the hair of a whitish grey. B. eapensis; M. eapensis, Gm. ; Taupe du Cap., Buff. Supp. VI. xxxvi. (The Rat-Mole of the Cape). Hardly as large as the guinea-pig; brown, with a spot round the ear, another round the eye, and a third on the vertex; the end of the muzzle white. The inci- sors are smooth. B. hottentottus, Less, and Garn., Voy. de la Coquille, pi. ii. (The Hottentot Rat-Mole). Smaller ; grey ; incisors smooth ; hardly as large as a rat. We must approximate to the Rat-Moles (Spalax and Bathiergus). Geomys, Rajin. — Pseudostoma, Say. — Ascomys, Lichten. The Geomys, which have four molars in compressed prisms throughout ; the first double, the remaining three simple ; the upper incisors furrowed with a double groove in front ; five toes to each foot ; the three middle anterior nails, that of the medius particularly, very long, crooked, and trenchant. They are low animals, and have very deep cheek-pouches, which open ex- ternally, enlarging the sides of the head and neck in a singular manner. One species only is known. G. bitrsarius; Mus bursarius, Shaw.* (The Canada Hamster). Size of a rat; fur of a reddish-grey; tail naked, and but half the length of the body. Inhabits deep burrows in the interior of North America. Diplostoma, Rafin. The Diplostomae are in every respect similar to the Geomys, except that they have no tail.-f- These animals are also from North America. The species before us is reddish, and ten inches in length. We now pass to larger Rodentise than those of winch we have hitherto * The figures of this animal, first published Trans. Lin. Soc. Vol. V. pi. viii, and Shaw, Vol. II. part 1, pi. 138, represent it with the internal skin of the cheek-pouches turned inside out, as though it had two sacs to the sides of the head. There is no- thing like it in nature. It is well represented, Acad. Berlin, 1S22 and 1823, pi. ii. f M. Rafinesque describes them as having only four toes to each foot. The Eu- ropean species has five, like the Geomys, RODENTIA. ]33 spoken, but of which several still have well developed clavicles. Of this number is the Castor, Lin. The Beavers are distinguished from all other Rodentia by their hori- zontally flattened tail, which is nearly of an oval form, and covered with scales. They have five toes to each foot : those of the hinder ones are connected by membranes, and that next to the thumb has a double and oblique nail. Their grinders, to the number of four throughout, and with flat crowns, appear as if formed of a bony ribbon reflected on itself, so as to shew one sloping edge at the internal border and three at the ex- ternal one of the upper row; in the lower ones it is exactly the reverse. Beavers are large animals, whose life is completely aquatic; their feet and tail aid them equally well in swimming. As their chief food is bark, and other hard substances, their incisors are very powerful, and grow as rapidly from the root as they are worn away at the point. With these teeth they cut trees of every description. Large glandular pouches which terminate on the prepuce produce a highly odorous oily substance, employed in medicine under the name of Castor {a). The organs of generation in both sexes terminate in the ex- tremity of the rectum, so that there is but a single external opening. C. fiber, Buff. VIII. xxxvi. (The Beaver). Larger than the badger, and of all quadrupeds the most industrious in constructing a dwelling, to effect which these animals act in concert in the most solitary parts of North America. Beavers choose water of sufficient depth to be frozen to the bot- tom, and as far as possible, running streams, in order that the wood which they cut above may be carried downwards by the current to the spot where it is to be used. They keep the water at an equal height, by dams composed of all sorts of branches, mixed with clay gSgT (a) The pouches of the beaver here alluded to are found in both sexes, being situated in the male behind the prepuce, and in the female at the upper edge of the orifice of the vagina, where they open. They are composed of dense cellular tissue, forming several folds, between which the castor is inclosed, and to which it adheres. The two pouches lie parallel with each other beneath the skin; they hang together, and separate a little at one of the extremities, which is larger and more rounded than the other. Their outer surface is smooth, of a dark brown colour, and free from hairs. The castor completely fills the pouches, but has a cavity in the centre, which is a distinguishing character that shews the genuine from the artificial article. When examined in the animal, the secretion is soft, being of a consistence somewhat in- termediate between wax and honey; but, when taken out, it dries, but does not be- come hard. The best castor used to come from Russia, in roundish, solid pods, smooth on the outside, and, when cut, presenting an orange-coloured surface. At present, the chief portion imported into this country is from Canada; it is brought in thin oblong pods, which are corrugated on the outside, and the castor contained in them is more deeply coloured than the Russian. Castor has a very peculiar, strong, and disagreeable odour; its taste is bitter, acrid, and slightly aromatic. Chemistry has discovered in castor a variety of substances, of which castorine is the principal. Castor is employed in medicine as a powerful antispasmodic in hysteric cases, and its effect is described by physiologists as being specifically applied to what are called the cerebro-spinal nerves, or those which have a mixed origin from the brain and spinal marrow. When taken, even to a small extent, it manifests its influence in the urine by its very peculiar flavour; but its efficacy as a remedy has considerably fallen in credit. — En